In firefox, it happens when saving a image.

As I noticed, not all the directory switching actions will trigger the
bug.  The current directory (location before you switches) should
contain some individual files of a certain type, then when switching
action happens, the filename is replaced by the first individual
file's name in that directory before disappears, which takes only .5
seconds or less.

I attached a video(screencast) in the duplicate report showing two
consecutive switching actions, of which only one triggers this bug.
But unfortunately the video can not catch the filename replacement
before disappearance.

The video could be found here:
http://librarian.launchpad.net/7044342/nautilus-save-as-bug.ogg

-- 
[Feisty] File save dialog delets filename when changing directory
https://launchpad.net/bugs/93396

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