I filed bug 949029 earlier and found that this happens only when
touchpad is being used, is the issue you are experiencing the same one?

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https://bugs.launchpad.net/bugs/949414

Title:
  menu items are not highlighted

Status in “gtk+3.0” package in Ubuntu:
  Confirmed
Status in “gtk+3.0” source package in Precise:
  Confirmed

Bug description:
  In GTK applications menu items are not always highlighted on mouse
  over (orange background).

  12.04 - 20120307

  When using touchpad, the menu items are not highlighted. When using a
  USB mouse or TrackPoint the widgets are reacting properly.

  STR:
  1. Open nautilus
  2. Open the "File" menu, OR right-click in the window, OR open any of the 
option menus inside the Preferences window.
  3. Move down through the menu using touchpad.
  4. Click on a sensitive menu item.

  Actual results:
  3. The first sensitive item is highlighted, but (usually) none of the other 
sensitive items are.
  4. The item is highlighted only once you click it.

  Expected results:
  Highlight should follow the mouse

  If global menu is disabled (env UBUNTU_MENUPROXY= ) , the menus do not
  react to the movement at all.

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