It sounds like your routing-group-connector isn't bi-directional.

Open the Exchange Management Shell, enter "get-routinggroupconnector" and make 
sure both sides are correct.

From: Lynden A. Philadelphia [mailto:lphiladelp...@philadelphiagroup.com]
Sent: Tuesday, December 22, 2009 9:12 PM
To: MS-Exchange Admin Issues
Subject: Exchange 2003 To 2007

Hello,

I have an 2003 Exchange environment that is working with no issues.

I recently installed Exchange 2007.

I forgot to suppress the Link State and now have done so.

I can send mail from a user on Exchange 2007 to the internet and to a user on 
2003.

I can't send mail from Exchange 2003 to 2007?

Can someone help me with this?

Help?

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