The three languages in the upper right are the last ones that were selected 
by the user.  The right language menu for destination language picks the 
output language for the next translation, which may replace one of the 
three int the upper right.

The source language selector is similar - it will show the recently used 
languages chosen by the left selector.

Try a translation, then change the output language and watch what happens. 
 You won't break anything ;)

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