Dear Viplav Ji,

I am sorry for limited available photographs in my thread of *Morinda*.
Though I disagree that those limited images do not carry enough info to id
my plant to species level, I accept your view that the heterostyly
character of this genus rejects the idea of determining a species level id
 based only on its stamen character.

Thank you very much for spending some time for me.

Regards

surajit




On Fri, Apr 24, 2015 at 11:07 AM, [email protected] <[email protected]>
wrote:

>
> Dear Surajit ji,
>
> The occurrence of distylous flowers in the genus *Morinda* has been
> documented in several sources including those quoted in my previous email.
> Once we admit to the existence of heterostyly in this genus, don't you
> think it negates the core basis upon which *M. exserta* was conceived and
> separated from the rest i.e. exsertion of the stamens and inclusion of the
> stigma? The possibility of the same occurring in the entire genus may have
> been either discounted or overlooked at the time.
>
> *Morinda exserta* Roxb. was described in the second volume of Roxburgh's
> Flora Indica, first published posthumously in 1824 (pp. 199-200
> <http://books.google.co.in/books?id=V1w-AAAAcAAJ&printsec=frontcover#v=onepage&q=morinda%20exserta&f=false>).
> Roxburgh would have recorded his observations much before his demise in
> 1815. In the year 1866, fifty-one years after Roxburgh's passing, "Heterostyly
> was first defined by Hildebrand (1866) as the presence of di- or
> trimorphism in the length of the style and stamens between flowers on
> separate plants of the same species.
> <http://onlinelibrary.wiley.com/doi/10.1111/j.1469-8137.1981.tb02348.x/pdf>
> "
>
> With the benefit of hindsight, we can sense the tenuous premise upon which *M.
> exserta* was based, at a time when thorough investigation into the
> phenomenon of heterostyly was decades away.
>
> It would be interesting to learn about any breakthrough involving the
> plant in your thread
> <https://groups.google.com/d/msg/indiantreepix/V7HGakB7kYQ/APFWz1ed2qMJ>.
> I think I have already conveyed my reluctance-cum-inability to attempt an
> identification of your individual plant from the limited available
> photographs.
>
> Best wishes, Viplav
>
> PS: The quote on heterostyly in the 2nd para. has been sourced from the
> journal New Phytologist, vol. 89, issue 4, p. 693, Dec. 1981. The PDF is
> hyperlinked in the quoted text.
>
> El 22 de abril de 2015, 22:17, surajitkoley <[email protected]>
> escribió:
>
>>
>> Thank you very much VIplav Ji for this new angle - heterostyly. Very
>> interesting and much probable solution of your two morphed population over
>> there. But it doesn't explain my query. It doesn't explain stipule
>> character. Neither it explains hair property of corolla, within as well as
>> without. As I said the flowers of my species may be a little hairy, but not
>> like what had been recorded in Cooke's Flora. Leaves are rather glabrous or
>> pubescent, not tomentose (Ref. Cooke's flora) specially underside, in my
>> species as far as I can make out examining my records.
>>
>> In short "heterostyly" doesn't explain if *Morinda tinctoria* var.
>> *tomentosa* and *Morinda exserta* is same species or not.
>>
>> I said Bangladesh document doesn't sastisfy my records, in addition to
>> what you have pointed out about the line drawing, the corolla size is
>> smaller in that doc., style is longer than corolla tube.
>>
>> Thank you very much once again.
>>
>> Regards
>>
>> surajit
>>
>>
>> On Wed, Apr 22, 2015 at 4:19 PM, [email protected] <[email protected]>
>> wrote:
>>
>>>
>>> Thank you Radha ji & Surajit ji for your thoughts. I am also intrigued
>>> by the reason behind different individuals of this tree bearing separate
>>> kinds of flowers i.e. with 'exserted anthers - included stigma' and vice
>>> versa. I have not come across these two different flower types/morphs on a
>>> single individual.
>>>
>>> Based on the above, would it be correct to infer that this tree is a
>>> distylous plant exhibiting heterostyly
>>> <http://en.wikipedia.org/wiki/Heterostyly>?
>>>
>>> In the description of the genus *Morinda* in Flora of China
>>> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=121192>,
>>> the flowers are said to be "bisexual and distylous".
>>>
>>> Flora of China
>>> <http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=121192>
>>> further states that "Morinda includes a notable range of breeding systems
>>> (Johansson, Opera Bot. 122: 1-167. 1994), but most of the species are
>>> apparently distylous, with the anthers and stigmas separated and their
>>> positions reciprocal between the short-styled and long-styled form of the
>>> same species; however, this biology has been sometimes overlooked."
>>>
>>> Best wishes, Viplav
>>>
>>> PS: In the paper on *Morinda* in Bangladesh, everything is exserted in
>>> the line drawing depicting the flower of *M. pubescens* i.e. anthers as
>>> well as stigma. I have not come across such a flower so far.
>>>
>>> El 21 de abril de 2015, 21:26, surajitkoley <
>>> [email protected]> escribió:
>>>
>>>>
>>>> I wonder Viplav Ji, if anybody ever examined those two version of
>>>> trees, bearing different types of flowers in the said locality, a bit more
>>>> closely!
>>>>
>>>> Does any version or even a single individual have two different types
>>>> of flowers on the same body? If yes what is the percentage? Are those two
>>>> types of flowers identical in other respect?
>>>>
>>>> If no, then they are likely to be two different taxa or var.
>>>>
>>>> It is interesting to note what Bangladesh
>>>> <http://www.banglajol.info/index.php/BJB/article/download/9766/7250>
>>>> think of related nomenclature. Please note that the flower description in
>>>> that doc doesn't tally with my record. You may check if it tallies with
>>>> yours. Also attached here what Haines recorded about *tomentosa*.
>>>>
>>>> Thank you
>>>> Regards
>>>> surajit
>>>>
>>>>
>>>>
>>>> On Tue, Apr 21, 2015 at 11:41 AM, [email protected] <[email protected]>
>>>> wrote:
>>>>
>>>>>
>>>>> Dear Surajit ji,
>>>>>
>>>>> It is difficult for me to say anything definitive from these
>>>>> photographs alone. Another problem, as mentioned in my previous email to
>>>>> Radha ji, is that there is substantial confusion surrounding the current
>>>>> nomenclature & synonymy of *Morinda* spp. (including *M. exserta *Roxb
>>>>> *.*).
>>>>>
>>>>> In my posts, I have used the name *Morinda coreia* var. *tomentosa* as
>>>>> per the keys by RR Fernandez
>>>>> <https://08511630493324166816.googlegroups.com/attach/83410cd801743ca3/08_Morinda_key_Fernandez.jpg?part=0.8&view=1&vt=ANaJVrHfmAvymogqKvW7zVddTOJt05S28mK4_Mr0jQVPWxf2n5DNr9akhVea9Q7w2dhbNACRYPnarBMPYXzMqaJ3OdGTbh9yw6h4G6tzQLq2zXZjG-VZhaA>
>>>>>  from
>>>>> Mumbai.
>>>>>
>>>>> Best wishes, Viplav
>>>>>
>>>>> El 20 de abril de 2015, 22:26, surajitkoley <
>>>>> [email protected]> escribió:
>>>>>
>>>>>>
>>>>>> Sorry, Viplav Ji, here are the photographs, cropped from the highest
>>>>>> resolution (not original) I have retained.
>>>>>>
>>>>>> Thank you
>>>>>> Regards
>>>>>> surajit
>>>>>>
>>>>>>
>>>>>> On Mon, Apr 20, 2015 at 10:24 PM, surajit koley <
>>>>>> [email protected]> wrote:
>>>>>>
>>>>>>> Thank you very much Viplav Ji for your kind feedback. Yes, the
>>>>>>> stipules are different in your thread (i had noted that. My species is 
>>>>>>> *M.
>>>>>>> exserta* Roxb., I think, which can be ascertained by its emarginate
>>>>>>> stipules and shape of leaf-blade (please check the top most leaf in the
>>>>>>> herb. -
>>>>>>> http://www.br.fgov.be/RESEARCH/COLLECTIONS/HERBARIUM/detail.php?ID=487591).
>>>>>>> I do not know if the corolla tube was glabrous inside & hairy outside.
>>>>>>> Perhaps a little hairy as can be seen in the pictures I attach herewith.
>>>>>>>
>>>>>>> Your species may well be different. I didn't mean that your species
>>>>>>> was *M. exserta*. In fact I did note the stipule as you have
>>>>>>> mentioned and also difference in leaf-lamina. I wanted your view on my
>>>>>>> thread as it was left unattended and other eFI uploads of *M.
>>>>>>> coreia* had flowers with inserted stamens.
>>>>>>>
>>>>>>> Thank you once again.
>>>>>>>
>>>>>>> Regards
>>>>>>>
>>>>>>> surajit
>>>>>>>
>>>>>>>
>>>>>>> On Mon, Apr 20, 2015 at 9:18 PM, [email protected] <
>>>>>>> [email protected]> wrote:
>>>>>>>
>>>>>>>>
>>>>>>>> Dear Surajit ji,
>>>>>>>>
>>>>>>>> Thank you very much for the link to your thread
>>>>>>>> <https://groups.google.com/forum/#!msg/indiantreepix/V7HGakB7kYQ/APFWz1ed2qMJ>.
>>>>>>>> The stamen exsertion in your flowers is similar to what I've posted. 
>>>>>>>> But
>>>>>>>> the stipules in your photographs appear to be larger, roundish &
>>>>>>>> persistent, quite unlike the small (c. 4 mm) triangular stipule
>>>>>>>> <https://groups.google.com/forum/#!topic/indiantreepix/ZV8KoQkdK6M> 
>>>>>>>> seen
>>>>>>>> by me which is caducous. In the flowers I've posted, the corolla
>>>>>>>> tube is glabrous on the inside and hairy on the outer surface
>>>>>>>> <https://08511630493324166816.googlegroups.com/attach/83410cd801743ca3/07_Morinda%20coreia%20var.%20tomentosa_flower.jpg?part=0.7&view=1&vt=ANaJVrFI-jRuhcmPsviwaDnfaYHTJ_SyDei2CS8KWHUXoXC-UwlObWFoDIn4Es8NT2fXP7dSj2Wy0neBO-ny1onS07yow5SLQ97uvzHARoh8ntwlnTqJp-Y>;
>>>>>>>>  I'm
>>>>>>>> unable to ascertain the same in your photographs. It would be 
>>>>>>>> interesting
>>>>>>>> also to see the fruits of your plant if possible.
>>>>>>>>
>>>>>>>> Best wishes, Viplav
>>>>>>>>
>>>>>>>> El 20 de abril de 2015, 7:05, surajitkoley <
>>>>>>>> [email protected]> escribió:
>>>>>>>>
>>>>>>>>>
>>>>>>>>> VIplab Ji,
>>>>>>>>>
>>>>>>>>> Can you please take a look at my thread
>>>>>>>>> <https://groups.google.com/d/msg/indiantreepix/V7HGakB7kYQ/APFWz1ed2qMJ>
>>>>>>>>> and give your view?
>>>>>>>>>
>>>>>>>>> Thank you
>>>>>>>>> Regards
>>>>>>>>> surajit
>>>>>>>>>
>>>>>>>>>
>>>>>>>>>
>>>>>>>>>
>>>>>>>>> On Sunday, 19 April 2015 10:41:48 UTC+5:30, Viplav Gangar wrote:
>>>>>>>>>>
>>>>>>>>>>
>>>>>>>>>> Sharing some photographs of the buds and the flowers of *Morinda
>>>>>>>>>> coreia* var. *tomentosa*. The flowers are usually 5-merous; have
>>>>>>>>>> also attached some instances of variable merosity in the flowers of 
>>>>>>>>>> this
>>>>>>>>>> tree (4-merous & 6-merous). The corolla tube is glabrous on the 
>>>>>>>>>> inside and
>>>>>>>>>> hairy on the outer surface as seen in the last picture.
>>>>>>>>>>
>>>>>>>>>> This tree is often referred to as *Morinda pubescens* Sm. The
>>>>>>>>>> genus *Morinda* in Mumbai was revised by RR Fernandez who
>>>>>>>>>> published a new combination - *Morinda coreia* Buch.-Ham. var.
>>>>>>>>>> *tomentosa* (Hook.f.) R.R.Fernandez. Have attached the key by
>>>>>>>>>> Fernandez from his book 'Trees of Mumbai (Bombay)', pp. 173-174, 
>>>>>>>>>> Scientific
>>>>>>>>>> Publishers, 1999.
>>>>>>>>>>
>>>>>>>>>> In Cooke's Flora, the anthers are said to be included within the
>>>>>>>>>> corolla tube as depicted in Navendu ji's photographs on
>>>>>>>>>> flowersofindia.net
>>>>>>>>>> <http://www.flowersofindia.net/catalog/slides/Indian%20Mulberry.html>.
>>>>>>>>>> But the anthers are clearly exserted in the trees that I have come 
>>>>>>>>>> across
>>>>>>>>>> in Madh. In the eflora archives, I found some photographs of the 
>>>>>>>>>> flowers of *M.
>>>>>>>>>> pubescens* bearing exserted stamens & anthers (posted by Mohina
>>>>>>>>>> ji fr. Alibag
>>>>>>>>>> <https://groups.google.com/forum/?hl=en#!topic/indiantreepix/qtNY1MByXak>
>>>>>>>>>> & Dinesh ji fr. SGNP in Mumbai
>>>>>>>>>> <https://groups.google.com/forum/?hl=en&fromgroups#!topicsearchin/indiantreepix/Morinda$20AND$20pubescens$20AND$20Sm./indiantreepix/mOj5Q4dogw8>).
>>>>>>>>>> The flower image of *M. pubescens* in 'The Trees of Mumbai' by
>>>>>>>>>> M. Almeida & N. Chaturvedi (p. 20) as well as in Shrikant ji's 
>>>>>>>>>> 'Trees of
>>>>>>>>>> Pune' (p. 98) also depicts exserted anthers. Would appreciate any
>>>>>>>>>> information / comments on this.
>>>>>>>>>>
>>>>>>>>>> Am sending the photographs of the leaves, fruit, and bark in
>>>>>>>>>> separate emails.
>>>>>>>>>>
>>>>>>>>>> Photographed on Madh Hill (North Mumbai) in March 2015.
>>>>>>>>>>
>>>>>>>>>> Best wishes, Viplav
>>>>>>>>>>
>>>>>>>>>
>>>>>>>>
>>>>>>>
>>>>>>
>>>>>
>>>>
>>>
>>
>

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