Boraginaceae is an interesting family of flowering plants often easily
recognised by somewhat rough stems and coiled inflorescence. Here is some
information to help members with uploads of members of this family. Members
are requested to participate liberally with uploads of both identified and
unidentified photographs.

Boraginaceae A. L. de Jussieu  Borage family
117 genera,  2,435 species
Widely distributed in temperate, tropical and subtropical regions.

*Salient features:* Bristly herbs, stems cylindrical, leaves alternate,
inflorescence helicoid cymes, flowers pentamerous, actinomorphic, carpels
2, ovary 4-lobed, style gynobasic, fruit with 4 nutlets.

Major genera:  Cordia  (300 species), Heliotropium (250) Tournefortia
(240), Onosma (140), Myosotis (90), Cynoglossum (75) and Ehretia (75).

Description: Herbs, shrubs or trees (Cordia), sometimes lianas, inner
phloem lacking, hairs with basal cystolith and often calcified or
silicified and as such plants bristly, rough to touch. Leaves alternate,
simple, entire, venation pinnate, reticulate, stipules absent.
Inflorescence usually of helicoid cymes, rarely scorpioid. Flowers
bisexual, actinomorphic, rarely zygomorphic (Echium), hypogynous,
pentamerous. Calyx with 5 sepals, free or slightly connate at base,
persistent. Corolla with 5 petals, united, rotate,  tubular or funnel
shaped, usually plicate. Androecium with 5 stamens, epipetalous, inserted
in corolla tube, filaments free, bithecous, anthers introrse, dehiscence
longitudinal, pollen grains tricolporate or multiporate, filaments often
with nectar discs at base. Gynoecium with 2 united carpels, ovary superior,
deeply 4-lobed, bilocular, axile placentation, becoming 4-locular due to
false septa, style 1, terminal or gynobasic, stigma 1 or bilobed, capitate
or truncate, ovary seated on a nectary. Fruit a drupe with one 4-seeded,
two 2-seeded or four 1-seeded pits or schizocarp with four 1-seeded
nutlets; seeds with embryo straight or curved. Pollination mostly by
insects. Drupaceous fruits are dispersed by birds, whereas corky ones
(Argusia, Cordia) are carried away by water.

Economic importance: Several species of Heliotropium (heliotrope),
Mertensia (virgin bluebells), Myosotis (forget-me-not), Cordia (Geiger
tree), Cynoglossum (hound’s tongue) and Pulmonaria (lungwort) are grown as
ornamentals. Several species  such as Borago officinalis (borage),
Symphytum officinalis (comphrey) and Lithospermum spp. (pucoon) have been
used as medicinal herbs. Alkanna tinctoria (alkanet) is a source of red dye
used to stain wood and marble and to colour medicines, wines and cosmetics.

Expected genera in India

*Adelocaryum*
*Anchusa*
Arnebia
Asperugo
Bothriospermum
Chionocharis
Cordia
Cynoglossum
Ehretia
Echinospermum
Eritrichium
Hackelia
Heliotropium
Lappula
Lindelofia
Lithospermum
Lycopsis
Maharanga
Mertensia
Microula
Myosotis
Onosma
Paracaryum
Pseudomertensia
Rochelia
Solenanthus
Trigonotis



-- 
Dr. Gurcharan Singh
Retired  Associate Professor
SGTB Khalsa College, University of Delhi, Delhi-110007
Res: 932 Anand Kunj, Vikas Puri, New Delhi-110018.
Phone: 011-25518297  Mob: 9810359089
http://www.gurcharanfamily.com/
http://people.du.ac.in/~singhg45/

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