Dear MARMAM colleagues,

We are pleased to announce that the following paper is now available online:

Enrico Pirotta, Peter L. Tyack, Jeffrey Adams, Michael J. Asaro, Phil J.
Bouchet, Daniel Crocker, John W. Durban, Ailsa J. Hall, Catriona M. Harris,
Amy R. Knowlton, Scott D. Kraus, Caroline Lehoux, Daniel W. Linden,
Caroline P. Good, Erin Meyer-Gutbrod, Alicia S. Miller, Carolyn A. Miller,
Patrick J. O. Miller, Michael J. Moore, Christopher D. Orphanides, Eric M.
Patterson, Heather M. Pettis, Theoni Photopoulou, Stéphane Plourde,
Nicholas R. Record, Jessica V. Redfern, Jason J. Roberts, Robert S. Schick,
Burton Shank, Laura Solinger, Brandon L. Southall, Marisa L. Trego, Len
Thomas. The combined effects of multiple stressors in an endangered,
long-lived species: Lessons learnt and ways forward. Ecological
Applications. A PDF copy of the paper can be downloaded for free from:
https://doi.org/10.1002/eap.70144

This paper is the result of a large collaborative effort to assess the
combined effects of multiple stressors on North Atlantic right whales. It
reports work conducted as part of one of the case studies within the
Population Consequences of Multiple Stressors (PCoMS) project. The
modelling approach presented here builds on methods described in two
previous papers, available at https://doi.org/10.1111/oik.09801 and
https://doi.org/10.1098/rsos.240050.

Abstract:
Exploring solutions to expanding industrial activities and climate change
requires assessments of the combined effects of multiple stressors on
wildlife populations. We present a spatially explicit state-space model for
the health, survival, reproduction, and somatic growth of individuals in a
long-lived, wide-ranging species. The model is applied to critically
endangered North Atlantic right whales (*Eubalaena glacialis*) to
investigate the combined effects of three primary stressors affecting the
species’ viability: entanglements in fishing gear, vessel strikes, and prey
availability. We estimate exposure to these stressors in space and time and
assess how their effects may combine in the pathway from exposure to vital
rates. Results suggested that changes in whale distribution after 2010 led
to increased entanglement risk. Poorer prey conditions were associated with
an increased effect of carrying fishing gear, but, overall, results on
combined effects were not conclusive and depended on model formulation. We
also incorporated the estimated effects of stressors into a population
viability analysis to explore alternative scenarios of stressor reduction.
This integrated analysis highlighted the importance of the declining trend
in maximum body length and its effect on reproduction, in addition to the
documented impact of entanglements on survival. Model development and
application elucidated critical data needs and the influence of underlying
mechanistic assumptions. Specifically, models for the combined effects of
stressors hinge on the availability of extended longitudinal measurements
of individual health and life history outcomes, extensive datasets on the
spatio-temporal distribution of stressors, and information on individual
space use affecting rates of exposure to stressors. Lessons from this
data-rich case study will support the generalisation of the modelling
approach to other long-lived species where measuring the population-level
consequences of multiple stressors directly is unfeasible.

Keywords: Bayesian state-space modelling; cumulative risk;
entanglements; *Eubalaena
glacialis*; North Atlantic right whale; population consequences of multiple
stressors; prey availability; stressor interactions; vessel strikes.

Please do not hesitate to contact me for any questions regarding our work.

Best Regards,
Enrico Pirotta
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