Dear MarMamers
We would like to share our new open access article that measures the metabolic 
contribution of digestion in the bottlenose dolphin 
(https://doi.org/10.1242/jeb.251474). This study quantified the heat increment 
of feeding (HIF), the metabolic cost of digestion, in eight common bottlenose 
dolphins using breath-by-breath respirometry. After consuming standardized fish 
meals (1659–2658 kcal), metabolic rate rose by ~37% above fasting levels and 
peaked ~60 min post-feeding, returning near baseline within two hours. Scaled 
to a full day, digestion increased daily metabolic needs by ~8.2% of basal 
metabolic rate, indicating a moderate energetic impact comparable to other 
marine mammals. These findings provide the first multi-individual estimate of 
HIF in dolphins, refining bioenergetic models and improving predictions of prey 
requirements and energetic vulnerability in wild populations

Title: Heat increment of feeding in the common bottlenose dolphin (Tursiops 
truncatus) contributes moderately to field metabolic rate estimates
Authors: Koliopoulou, I., DeRuiter, S.L. Altimiras, J., Larsson, J., Arenarez, 
J., Rosen, D.  Fahlman, A.
doi: https://doi.org/10.1242/jeb.251474
Abstract: Digestion elevates metabolism through the heat increment of feeding 
(HIF) – the energy expended on mechanical and biochemical processes after 
eating. Quantifying this cost is essential for bioenergetic models that predict 
energy flow and prey requirements in populations. Using breath-by-breath 
respirometry, we measured oxygen consumption (V̇ O2) in eight common bottlenose 
dolphins (Tursiops truncatus) before and after feeding standardized meals 
(1659–2658 kcal of capelin and herring). Metabolic rate rose by ∼37% above 
resting levels, peaking 60 min after feeding before returning to baseline 
within 2 h. When scaled across the day, digestion increased daily metabolic 
needs by ∼8.2% of basal metabolism, similar to values reported for Steller sea 
lions (Eumetopias jubatus) and harbour seals (Phoca vitulina), where HIF 
contributes 4–10% of daily energy expenditure. This study provides the first 
multi-individual estimate of HIF in dolphins and suggests that the energetic 
cost of digestion is a moderate contribution to overall daily metabolism, 
refining energetic models and improving prey requirement estimates for 
cetaceans in the wild.
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