On Thu, 17 Apr 2014 21:09:05 -0400 Meade Dillon <dillonm...@gmail.com>
wrote:

> Squirrels are VERY important.  They open pine cones and bury the seeds,
> bury acorns, walnuts, butternut, etc.  Without squirrels, we'd have no
> forests.  Some misguided souls think forest fires open pine cones which
> allow new pine trees to sprout and grow, but that is an old wives
> tale...

Actually, it seems it is not an old wives tale.
----------------------------
http://www.nps.gov/fire/wildland-fire/learning-center/educator-resources/lesson-plans/fire-adaptation.cfm

Retention of seeds by plants and stimulation of seed dispersal by fire
are other examples of fire-adaptive strategies. A number of pine species
have pine cones that open only after a fire. These cones are said to be
serotinous (pronounced sir-ot-in-ous). Jack pines have cones that are
held closed by a resin that is sensitive to high temperatures. These
cones will not open to release their seeds until the critical temperature
is reached. Lodgepole pine cones (a western U.S. variety of tree) vary
from serotinous to free-opening. When these trees grow in areas subject
to frequent fires, the cones are serotinous. However, if this species
grows in areas where fire is less frequent, the cones open and release
their seeds more frequently without fire.
----------------------------
http://interwork.sdsu.edu/fire/resources/conifer-forest.htm

Coulter pine displays variation in degree of cone serotiny (cones that
open with fire). Not every Coulter pine cone is serotinous.  There are
some trees with no serotinous cones and some with all serotinous cones
and everything in between.  Serotiny is prevalent in Coulter pines
occurring withing chaparral or oak woodland.  Cones of Coulter pine in
these communities typically do not open until heated by fire.
Consequently, the bulk of Coulter pine regeneration in these communities
occurs after fire.  Coulter pine ecotypes associated with coast live oak,
however, typically bear cones that open at maturity or shortly
thereafter.  Coulter pine seedling development is best in mineral soil in
open areas; such conditions are created by fire.
----------------------------
http://indianapublicmedia.org/amomentofscience/lodgepole-pine-trees-love-forest-fires/

The summer of 2012 was certainly a scorcher, with forest fires blazing
across the Western United States. To humans, wildfires are often scary
and dangerous. If you were a lodgepole pine cone, though, you would be
eagerly awaiting the blazing heat of those flames!

How come lodgepole pine cones like fire so much?

When lodgepole pines grow, especially in areas that are prone to forest
fires, their cones are tightly sealed. A layer of resin and woody tissue
sticks the cones’ scales together. The seeds are locked in tight, and the
cones can’t open unless they’re exposed to VERY high temperatures–the
type of temperatures that fire provides.

“Serotinous” is a scientific term for a seed that requires an
environmental trigger in order to be released. For the lodgepole pine,
that trigger is heat. And since big fires don’t come along very often,
those well sealed pine cones have to be extremely patient. They can hang
out on the tree branches for several years waiting for enough heat to
open them up!

Years of waiting

So, if multiple years’ worth of cones can accumulate, then a lot of new
pine trees sure must sprout up after a fire.

Lodgepole pines are famous for colonizing post fire landscapes. The seeds
love the carbon rich soil that fire leaves behind, and seedlings pop up
almost immediately. They grow into dense stands of trees, and before you
know it, there’s a whole new crop of serotinous cones waiting in
anticipation for the next fire to blaze through.
----------------------------
http://techalive.mtu.edu/meec/module11/FireandJackPine.htm

Why Do Jack Pine Trees Grow After Fire?

Michigan Environmental Education Curriculum
Forest Ecosystem

New jack pine seedlings grow well after a hot fire. Fire causes the cone
of an older jack pine tree to open up and release seeds. Cones that need
heat, such as the heat from fire, in order to open are called serotinous
cones. When the heat causes the cones to open, seeds are released and
fall to the ground (see animation below).

In Michigan, the Department of Natural Resources (DNR) lights fires in
the Jack Pine Ecosystems to help new jack pine trees grow. This is
because the Kirtland's Warbler will only build nests under the young jack
pine trees.
----------------------------
csfs.colostate.edu/pdfs/06307.pdf

Lodgepole Pine

Lodgepole pine forests are very adapted to a natural fire regime.
Regeneration of new lodgepole seedlings can be rapid (as little as two to
three years).

Lodgepole bark is thin in comparison to that of ponderosa pine.
Temperatures lethal to the cambium are common.

A unique characteristic of lodgepole pine is its serotinous cones. The
cone scales can remain closed for several years because of a resin
coating. During an intense fire this resin melts away allowing the cone
scales to open, thus releasing the seed. After a fire, a massive number
of seeds are released. An intense fire also exposes mineral soil to
provide a good seed bed.
----------------------------
http://michigan.gov/dnr/0,1607,7-153-10370_12146_12212-60297--,00.html

Jack Pine (Pinus banksiana)

Among all Michigan tree species, the jack pine is uniquely adapted to
exist and reproduce on the hottest and driest sites in Michigan. It
thrives on dune sand and on the sandy glacial plains, where it often
occurs in dense stands. It is called a "fire species" because wildfires
sweep through jack pine stands, killing the trees and preparing the
ground for a new stand, thus releasing seeds from its cones.

Jack pine was once thought to poison the ground on which it stood because
no other trees and few ground plants would grow near it. In truth, many
species grow with jack pine.

Most jack pine cones are sealed with a special resin. This resin prevents
the cones from drying out and releasing their seeds except under certain
conditions. The resin melts at 112 degrees F, a temperature easily
reached during a forest fire, but which also may be achieved on bare
ground in open sunlight on a bright summer day. After the resin is
melted, the cones open, releasing the seeds from within. These resinous
cones are called serotinous cones.

Not all jack pine cones are serotinous. Almost every jack pine has a few
cones each year which are non serotinous. These cones open in the fall
and provide a source of seeds in case fires do not occur.

Given proper conditions, the seeds will germinate almost immediately
after falling, but if moisture is lacking when they fall, some seeds go
into dormancy which may last up to three years or more. Studies have
shown jack pine seedlings will continue to sprout up to three years after
a forest fire. This adaptation guards against the loss of all of the
young seedlings due to an unusually dry summer or because of a hard frost.

Jack pines in Michigan are often crooked and short generally not a
favorite tree of most people. However, in some places in the Upper
Peninsula and farther north into Ontario, they grow much straighter and
taller. Much of the difference is due to insects and other problems that
plague the jack pine in lower Michigan, which is the southern limit of
the tree's natural range.

The long wood fibers of jack pine make it ideal for the manufacture of
strong papers. Harvest is usually by clearcut to mimic the effects of a
wildfire. The soil is then prepared for seeding or planting, and a new
stand is established.

Jack pine pollen is shed in late May or early June and sometimes forms
large clouds resembling smoke rising from the tops of the trees. It
settles on the ground and forms a yellow border around puddles and a
thin, yellow film on everything within miles.
----------------------------
http://www.fs.fed.us/database/feis/plants/tree/pinser/all.html

 REGENERATION PROCESSES:
The specific epithet of pond pine, serotina, means "late" and refers to
the delayed opening of cones. Cones often persist for 2 to 8 years,
giving the tree the appearance of being a prolific seed producer [8,14].
All pond pine cones are probably closed when they first mature [79].
Viability does not decrease in cones that remain closed for as long as 3
years [8]. Cones older than 5 years may open individually following
weathering and insect depredations; sound seed may be sparse in cones
more than 5 years old [79]. Seed is released from older, open cones of
pond pine during 2 seasonal periods: April through September  and October
through January [8,16].  Regeneration of pond pine is hampered by 2
problems under average conditions. First, serotinous pond pine cones
often require heat to open the scaled cones, and 2nd, continual moistness
of the organic soil provides an excellent growth medium for intense
competitors such as other trees, evergreen brush, and switchcane
(Arundinaria gigantea ssp. tecta) [16].

Pond pine cones open and release seeds soon after exposure to heat from
fire. Intensity of heat does not adversely reduce viability of seeds.
Even badly charred cones release seeds that are capable of germination.
Mature cones can be opened by exposure to  333 to 336 oF (167-169 oC) dry
heat for 20 seconds or by immersion in boiling water for a similar period
[8]. Optimum stratification of pond pine seeds is 30 days at 33-41oF
(0.5-5 oC) in a moist medium [5,46]. Germination occurs in 24 days [5].
Pond pine seeds germinated in a nursery were unable to survive full
saturation conditions [46].
----------------------------
http://oscodaherald.com/stories/Forest-Service-to-conduct-prescribed-burns-in-Oscoda-County,99161

[U.S. Forest Service Public Relations Manager Kate] Salm said in some
cases where fire is used to aid wildlife, it can have a huge impact for
plants and creatures.

According to Forest Service biologists, fire is a natural part of many
ecosystems in the Huron-Manistee National Forests. Species such as
white-tailed deer, turkey, butterflies, grouse and turtles use burned
areas for food, cover and a place to raise their young.

Generating new habitat for the endangered Kirtland’s warbler, a bird that
is largely only found in Michigan’s juvenile jack pine forests, is
another reason for the burns, Salm said.

A surprising reason, Salm said, is to create habitat for the monarch
butterfly, which she said has been on the decline over the past years.

The new fire areas create habitat for the monarch’s favorite flowering
plant, the milkweed, to grow, she said.

Another motive behind for prescribed burns is to allow native trees, like
jack pine and red pine, to release seeds.

“The trees will drop their cones on the ground; they need that heat from
the fire to open, so the seeds will spread,” she said. “That is something
with that tree that has developed over thousands of years.”

The fires also put large amounts of nutrients back into the soil, causing
new plant life to flourish.

“It doesn’t take long after a fire,” she said. “You see how green the
grass is after a prescribed fire. The ash puts a lot of nutrients back
into the soil.”

Salm said fire has been a natural process with Michigan’s ecosystem since
before humans were even in Michigan.

“Before Europeans most of it was caused by nature, often by lightning
strikes,” she said.
----------------------------

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