If your proxy is based on the openlayers example proxy.cgi, you will see 
that the error for a host that is not allowed is:

print "Status: 502 Bad Gateway"
print "Content-Type: text/plain"
print
print "This proxy does not allow you to access that location (%s)." % 
(host,)
print

This pretty much describe what you are seeing. I would say that your host
castle.iplantcollaborative.org 
<http://castle.iplantcollaborative.org:8080/geoserver/wms/?LAYERS=opengeo%3APimaPine,opengeo%3ABIEN_Species_Richness_Margalef,opengeo%3ABIEN_Species_Richness,opengeo%3ABIEN_Species_Richness_Margalef_Interpolated&QUERY_LAYERS=opengeo%3APimaPine,opengeo%3ABIEN_Species_Richness_Margalef,opengeo%3ABIEN_Species_Richness,opengeo%3ABIEN_Species_Richness_Margalef_Interpolated&STYLES=,,,&SERVICE=WMS&VERSION=1.1.1&REQUEST=GetFeatureInfo&BBOX=-18295967.087793%2C-10684061.894102%2C18295967.087793%2C10684062.234102&FEATURE_COUNT=10&HEIGHT=546&WIDTH=935&FORMAT=image%2Fpng&INFO_FORMAT=application%2Fvnd.ogc.gml&SRS=EPSG%3A900913&X=174&Y=140>
is not in the allowedHosts list or that you have misspelled it there.

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