I found the problem.
When I updated the template I uninstalled Thunderbird, and the qubes-usb-proxy 
and other related tools are considered unused dependencies. I think that this 
is a bug in the template. I solved the issue installing the necessary packages.

$ sudo dnf remove thunderbird
Dependencies resolved.
==============================================================================================
 Package                  Arch   Version         Repository                     
         Size
==============================================================================================
Removing:
 thunderbird              x86_64 52.4.0-2.fc26   @updates                       
        157 M
Removing dependent packages:
 python-backports-ssl_match_hostname
                          noarch 3.5.0.1-4.fc26  @fedora                        
         18 k
 python2-crypto           x86_64 2.6.1-14.fc26   @fedora                        
        2.2 M
 python2-cryptography     x86_64 2.0.2-2.fc26    @updates                       
        2.6 M
 python2-jinja2           noarch 2.9.6-1.fc26    @fedora                        
        3.6 M
 python2-pyOpenSSL        noarch 16.2.0-6.fc26   @updates                       
        523 k
 python2-requests         noarch 2.13.0-1.fc26   @fedora                        
        409 k
 python2-tornado          x86_64 4.4.2-3.fc26    @fedora                        
        3.9 M
 python2-urllib3          noarch 1.20-1.fc26     @fedora                        
        536 k
 python2-zmq              x86_64 16.0.2-3.fc26   @fedora                        
        1.6 M
 qubes-mgmt-salt-vm-connector
                          noarch 3.2.5-1.fc26    
@qubes-builder-vm-r3.2-current-testing 4.7 k
 qubes-vm-recommended     noarch 3.2.3-1.fc26    
@qubes-builder-vm-r3.2-current-testing   0  
 salt                     noarch 2017.7.2-1.fc26 @updates                       
         36 M
 salt-ssh                 noarch 2017.7.2-1.fc26 @updates                       
        3.0 k
 thunderbird-qubes        x86_64 1.2.9-1.fc26    
@qubes-builder-vm-r3.2-current-testing  11 k
 zeromq                   x86_64 4.1.6-3.fc26    @fedora                        
        671 k
Removing unused dependencies:
 PyYAML                   x86_64 3.12-3.fc26     @fedora                        
        646 k
 dnf-utils                noarch 2.1.5-1.fc26    @updates                       
        3.8 k
 libtomcrypt              x86_64 1.17-31.20160123git912eff4.fc26
                                                 @updates                       
        824 k
 libtommath               x86_64 1.0-8.fc26      @updates                       
        106 k
 libyaml                  x86_64 0.1.7-2.fc26    @fedora                        
        121 k
 openpgm                  x86_64 5.2.122-8.fc26  @fedora                        
        317 k
 python-backports         x86_64 1.0-9.fc26      @fedora                        
        638  
 python-chardet           noarch 2.3.0-3.fc26    @fedora                        
        1.1 M
 python-enum34            noarch 1.1.6-1.fc26    @fedora                        
        232 k
 python-ipaddress         noarch 1.0.16-4.fc26   @fedora                        
        227 k
 python2-asn1crypto       noarch 0.23.0-1.fc26   @updates                       
        924 k
 python2-babel            noarch 2.3.4-5.fc26    @fedora                        
         20 M
 python2-backports_abc    noarch 0.5-3.fc26      @fedora                        
         30 k
 python2-futures          noarch 3.1.1-1.fc26    @fedora                        
         91 k
 python2-idna             noarch 2.5-1.fc26      @fedora                        
        563 k
 python2-markupsafe       x86_64 0.23-13.fc26    @fedora                        
         83 k
 python2-msgpack          x86_64 0.4.8-3.fc26    @fedora                        
        272 k
 python2-pysocks          noarch 1.6.7-1.fc26    @updates                       
         79 k
 python2-singledispatch   noarch 3.4.0.3-7.fc26  @fedora                        
         46 k
 python2-systemd          x86_64 234-1.fc26      @fedora                        
        262 k
 pytz                     noarch 2016.10-4.fc26  @fedora                        
        185 k
 qubes-core-vm-doc        noarch 3.2.20-1        
@qubes-builder-vm-r3.2-current-testing 2.2 k
 qubes-gpg-split          x86_64 2.0.26-1.fc26   
@qubes-builder-vm-r3.2-current-testing  59 k
 qubes-img-converter      x86_64 1.2.3-1.fc26    
@qubes-builder-vm-r3.2-current-testing 6.5 k
 qubes-input-proxy-sender x86_64 1.0.10-1.fc26   
@qubes-builder-vm-r3.2-current-testing  18 k
 qubes-pdf-converter      x86_64 2.1.2-1.fc26    
@qubes-builder-vm-r3.2-current-testing  15 k
 qubes-usb-proxy          noarch 1.0.11-1.fc26   
@qubes-builder-vm-r3.2-current-testing 7.7 k

Transaction Summary
==============================================================================================
Remove  43 Packages

Freed space: 235 M
Is this ok [y/N]:

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