Dear Julien:

I tried the attached Rscript with mvMORPH to attain a phylogenetic correlation matrix betwen all pairs of variables O - AQ in the attached csv file (variables B-N are just variables that define the assignment to various subgroups and won't be included in these analyses). The tree is also attached.

Howver, when I am running your script under these data, I am always getting various error messages. Maybe, I need to sort the species lists prior to the analysis, so that both data frame and tree show the same sequence, but the problem might be something else.

Would you mind to have a short check on my data?

Thanks a lot,

Oliver


Zitat von Julien Clavel <julien.cla...@hotmail.fr>:

Dear Oliver, 
 
Yes, you can use the independent contrasts to estimate the
correlations (e.g., assuming BM). Otherwise, you can compute the correlations
directly from the models fit in mvMORPH. 
 
For instance, under BM you can use: 
 
fit_bm <- mvBM(tree, data) 
cov2cor(fit_bm$sigma) # marginal correlations 
cor2pcor(fit_bm$sigma) # partial correlations 
 
Under OU, you can also retrieve the correlations from the variance-covariance matrix (which depends on both the “sigma” and “alpha” parameters, and can be retrieved
with the “stationary” function): 
fit_ou <- mvOU(tree, data) 
cov2cor(stationary(fit_ou$sigma)) # marginal correlations 
cor2pcor(stationary(fit_ou)) # partial correlations 
 
 
Alternatively, you can use the “mvgls” function to do it, e.g.: 

fit_bm2 <- mvgls(data~1, tree=tree, model="BM", method="LL") 
cov2cor(fit_bm2$sigma$Pinv) # marginal correlations 
cor2pcor(fit_bm2$sigma$Pinv) # partial correlations 
 
You can for instance use penalized likelihood to obtain a
regularized estimate of the evolutionary correlations: 
 
fit_bm2 <- mvgls(data~1, tree=tree, model="BM", method="PL") # Ridge penalization by default
cov2cor(fit_bm2$sigma$Pinv) # marginal correlations 
cor2pcor(fit_bm2$sigma$Pinv) # partial correlations 
 
With LASSO penalization, for instance, you can find a sparse
estimate for the partial correlations. That is, you can directly select the “significant”
partial correlations from the model fit: 
 
fit_bm2 <- mvgls(data~1, tree=tree, model="BM", method="PL",
penalty="LASSO") 
cov2cor(fit_bm2$sigma$Pinv) # marginal correlations 
cor2pcor(fit_bm2$sigma$Pinv) # partial correlations 
 
Best wishes, 
 
Julien 



De : R-sig-phylo <r-sig-phylo-boun...@r-project.org> de la part de Oliver Betz <oliver.b...@uni-tuebingen.de>
Envoyé : dimanche 13 juin 2021 14:45
À : r-sig-phylo@r-project.org <r-sig-phylo@r-project.org>
Objet : [R-sig-phylo] phylogenetic correlation analysis
 
Dear all:

I would like to perform a phylogenetic correlation analysis (simlar to 
PGLS, but correlation instead of regression), so that I get a 
correlation matrix, where all the Pearson or Spearman correlation 
coefficients between all of my variables are listed. One solution 
might be to calculate PICs and do standard correlation analyses on 
them, but there might be a more direct solution available?

Which R package would you recommend for such analysis?


Thank you very much,

Oliver Betz

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#Correlation analysis
#Data management to prepare correct data and tree file


library(mvMORPH)



setwd("c:/temp")
tree <- read.tree("DataR/transmitting science course/Own 
data/Steninae_reduced.nwk")


# load eye protrusion versus attack distance towards large springtails
data <- read.csv2("DataR/transmitting science course/Own data/Daten Mittelwert 
für Hauptkomponentenanalyse_Gesamtliste_Original April 2021_data 
transformed_nur Relativwerte_Phyl.csv", header=T)
names(data) <- c("Species","Gattung","Artengruppe", "Untergattung", 
"Klebpolstertyp", 
"Tarsustyp","Hoehe_1","Hoehe_2","Hoehe_3","Hoehe_4","Litter","Rocks","Vegetation","Ground","log_Körperlänge","log_Vorderkörper","log_Pronotuml","log_rel_Hinterbeinl","log_rel_Antennenl",
                 
"log_rel_Labiuml","LOG_eye_protrusion","log_rel_Haftpolsterfl","log_rel_Anz_Hafth_Klebp","LOG_rel_Anzahl_Haftkont_Klebp","LOG_rel_Breite_4.Vordertarsus",
 "LOG_rel_Anzahl_Hafthaare_4.Vordertarsus",
                 
"LOG_rel_Attachment_force_smooth","LOG_rel_Attachment_force_rough", 
"LOG_rel_attack_dist_small_Md", 
                 "LOG_rel_attack_dist_large_Md", 
"LOG_rel_attack_dist_small_Lb", "LOG_rel_attack_dist_large_Lb", 
"ARCSIN_capture_succ_Md_small", "ARCSIN_capture_succ_Md_large", 
"ARCSIN_capture_succ_Lb_small",
                 "ARCSIN_capture_succ_Lb_large", 
"Niche_width_elevation","Niche_width_habitat", "Species_freq_Doi_Inth", 
"Species_freq_Doi_Php",
                 "ARCSIN_selection_index_vertical", 
"ARCSIN_selection_index_ground", "ARCSIN_selection_index_stones") # rename 
colums with shorter names, save dataframe as "dat"
rownames(data) <- data$Species          # use species names to set rownames
#_________________________________________________

fit_bm <- mvBM(tree, data, model = "BM1") 
cov2cor(fit_bm$sigma) # marginal correlations 
cor2pcor(fit_bm$sigma) # partial correlations 
Species;Gattung;Artengruppe;Untergattung;Klebpolstertyp;Tarsustyp;Hoehe_1;Hoehe_2;Hoehe_3;Hoehe_4;Litter;Rocks;Vegetation;Ground;log_Körperlänge;log_Vorderkörperl;log_Pronotuml;LOG_rel_Hinterbeinl;LOG_rel_Antennenl;LOG_rel_Labiuml;LOG_eye_protrusion;LOG_rel_Haftpolsterfl;LOG_rel_Anzahl_Hafth_Klebp;LOG_rel_Anzahl_Haftkont_Klebp;LOG_rel_Breite_4.Vordertarsus;LOG_rel_Anzahl_Hafthaare_4.Vordertarsus;LOG_rel_Attachment_force_smooth;LOG_rel_Attachment_force_rough;LOG_rel_attack_dist_small_Md;LOG_rel_attack_dist_large_Md;LOG_rel_attack_dist_small_Lb;LOG_rel_attack_dist_large_Lb;ARCSIN_capture_succ_Md_small;ARCSIN_capture_succ_Md_large;ARCSIN_capture_succ_Lb_small;ARCSIN_capture_succ_Lb_large;Niche_width_elevation;Niche_width_habitat;Species_freq_Doi_Inth;Species_freq_Doi_Php;ARCSIN_selection_index_vertical;ARCSIN_selection_index_ground;ARCSIN_selection_index_stones
D.andrewesi;1;2; 
;1;2;1;1;0;0;0;1;0;0;,830588668685144;,527468782530814;,309630167425899;,642407442571537;,363878541576991;,116152348313469;,317854489331469;
 ; ; 
;,0336209064093937;,897109673499622;1,20682587603185;1,66558099101795;,280508328519379;,424033067340483;
 ; ;,570199503280313;0; ; ;,17;0;,12; ; ; ; 
D.betzi;1;1; 
;1;1;1;1;0;1;1;1;1;1;,66254063013571;,369345755134671;,240399465738641;,554005731593748;,225172209272876;,107485819173744;,443716607769532;
 ; ; 
;,0244340161199135;,936221491181442;2,01494034979294;2,05804623039528;,389614165156824;,295592378040602;
 ; ;,656060590924923;,477995198518952; ; 
;,066;,11;,15;,15;,00200000133333573;,144502365941527;1,02362572892893
D.chetri;1;28; 
;1;1;1;1;0;0;0;1;0;0;,870403905279027;,619441183654473;,341764159874347;,66838591669;,352316912196267;,152288344383056;,23946632295603;
 ; ; 
;,0424724041091961;,700346372889992;1,01703333929878;1,96473092105363;,114222431987793;,452311902925352;
 ; ;,635276383213721;0; ; ;,37;0;,09; ; ; ; 
D.fornicifrons;1;28; 
;1;2;1;0;0;0;1;1;0;0;,841234295506041;,599773939146388;,328950132069784;,673020907128896;,389166084364532;,115918674280109;,217299658976496;
 ; ; 
;,0464188947015681;,952039522330594;1,14921911265538;1,78247262416629;,157981073122323;,315952565084685;
 ; ;,617624310253785;,317823703927881; ; 
;0;,47;,09;,08;,0140004573736748;1,40326848329556;0
D.karen;1;2; 
;1;1;1;1;0;0;0;1;1;0;,704750904290671;,417969642214737;,253580289562183;,57287160220048;,330413773349191;,12057393120585;,289053557592641;
 ; ; 
;,0364489893366717;,935587552592184;1,87448181769947;2,73663549768682;,25848569872926;,261795198153571;
 ; ;,20064353830785;,101273024409407; ; 
;,4;,29;,27;,23;,00500002083356771;,412608195642369;,637265648700304
D.luteolunatus;1;29; 
;1;2;1;1;0;0;1;1;0;0;,780965029608317;,475089803389007;,276461804173244;,571708831808688;,309630167425899;,12057393120585;,311011748981288;
 ; ; 
;,035926420251516;,815651321782137;1,08635983067475;1,89652621748956;,198286317596186;,29190738046508;
 ; ;,40639442908634;,234338894401164; ; 
;,47;,13;,24;,23;,00100000016666674;,210552264721877;,912441752748123
D.meo;1;1; 
;1;1;1;1;0;0;0;1;0;1;,638589083292717;,353339095311305;,227886704613674;,584331224367531;,296665190261531;,0969100130080564;,445682026852679;
 ; ; 
;,0340420429872385;,917882192744799;1,64443858946784;1,99431715266964;,228593752039125;,316486946727806;
 ; ;,843086960339661;,173874786487402; ; 
;,43;,38;,09;,08;,00100000016666674;,205442117510134;,919007579501776
D.nitidicollis;1;28; 
;1;2;1;1;1;0;1;1;0;0;,852654234153498;,610964184440695;,339593390803342;,716837723299524;,531478917042255;,136720567156407;,230266406359028;
 ; ; 
;,0462096277662557;,697899073446361;,968482948553935;1,6222140229663;,284471504374667;,371890916572443;
 ; ;,430912433957024;,203705913292907; ; ;,47;,55;,15;,23; ; ; 
D.obliquenotatus;1;28; 
;1;2;1;1;0;0;0;1;0;1;,872331121230251;,630732892817196;,338854746252323;,684626674476734;,404503778174426;,135215190430548;,249589657772943;
 ; ; 
;,0340062524460803;,781513275388627;1,68663626926229;2,24772783290972;,331017713245311;,366557263298232;
 ; ;,388715439357427;,366017949297474; ; 
;,46;,17;,3;,15;,00400001066674347;1,49331728187102;0
D.pseudacutus;1;29; 
;1;3;0;1;0;0;0;1;0;0;,692229835772756;,462697408101717;,283074974735472;,617000341120899;,378397900948138;,117249650556733;,27214341759105;
 ; ; 
;,037919023916281;,914995733933636;2,02775720469055;2,58194965837332;,336191787314376;,36952675560824;
 ; ;,448484047723639;,338105027201183; ; 
;0;0;,03;,08;,0380091512810838;,986317927412527;,128352127729054
D.punctiventris;1;28; 
;1;2;1;0;0;0;0;1;0;0;,880870732532424;,574841195063385;,342027688087472;,606381365110605;,262451089730429;,126324055793894;,240649074620679;
 ; ; ;,0296947978482294;,759421208085003; ; ;,159589285075503;,242727209249022; 
; ;,366125045404757;,237218568094435; ; ;0;,12;,09; 
;,0150005625569608;1,39737400569929;0
D.scabricollis;1;28; 
;1;1;1;1;0;0;1;1;0;0;,723181928261923;,455859567203536;,273001272063738;,627950765074281;,382959950139757;,147543829864141;,269785265754435;
 ; ; 
;,0306552267004634;,83412320846046;1,22271647114758;1,76342799356294;,155249256671107;,191211403710181;
 ; ;,811905938993222; ; ; ;,3;,19;,09;,08; ; ; 
D.siamensis;1;30; ;1;3; ; ; ; ; ; ; ; 
;,649334858712142;,394451680826216;,227886704613674;,648289284739931;,423601998993174;,119054966543886;,339650157613684;
 ; ; ;,0639802413612563;1,04286490604421;2,20194306340165;2,13513265137677; ; ; 
; ; ; ; ; ; ; ; ;,08; ; ; 
D.siwalikensis;1;28; 
;1;1;1;0;0;0;0;1;0;1;,87633332908638;,629205657102304;,362293937964231;,635483746814912;,338456493604605;,113943352306837;,230065551206047;
 ; ; 
;,0299943009666998;,730857451151487;1,11727129565576;1,60852603357719;,234527383501801;,32972481522594;
 ; ;,609063977170518;,261055051689916; ; ;0;,17;,15; ; ; ; 
D.srivichaii;1;28; 
;1;2;1;1;1;0;1;1;1;0;,719993826367604;,470998169660874;,253822438708073;,661812685537261;,421603926869831;,15836249209525;,264794219407155;
 ; ; ;,0403563496555686;,879907418931813;1,64246452024212;2,59769518592551; ; ; 
; ; ; ; ; ;,51;,091;,45;,23;,0150005625569608;1,39737400569929;0
D.tortuosus;1;31; 
;1;2;1;1;0;0;0;1;0;1;,809559714635268;,525476726000246;,296299468570948;,606381365110605;,330413773349191;,12057393120585;,295786940251609;
 ; ; 
;,0314869252496651;,824229930737481;1,40823996531185;1,84198480459011;,179872667590043;,329520882212919;
 ; ;,716699831265603;,131175869380332; ; ;,17;,098;,09; 
;,00300000450001823;,276510115512615;,809583593564425
D.tubericollis;1;1; 
;1;1;1;1;0;0;0;1;0;0;,557958104678714;,302114376956201;,222716471147583;,517195897949974;,209515014542631;,0999004309846666;,448300428150309;
 ; ; 
;,0251009610468134;,840167747053471;1,85733249643127;2,09412159584056;,197589115496967;,339762821989302;
 ; ;,72856550462205;,157854757786628; ; 
;,3;0;,03;,15;0;,374707134712372;,686714729593209
S.amoenus;2;9;2;2;3;1;1;0;0;1;1;1;1;,764791280484541;,43756560486588;,279223943904862;,547774705387823;,227886704613674;,599883072073688;,54139961949684;,0269985442399249;1,23711637529876;1,87528853603335;,0428105819284757;1,10703632142884;1,40654018043396;1,76715586608218;,261737849803336;,220004006354273;,434296025514549;,49162198147494;,577343568411807;,167481885486474;,793910184547671;,357357607953973;,5;,86;,12;,23;,0590342835640201;,0430132622044499;1,11520271518489
S.angusticollis;2;9;2;2;3;1;1;0;0;1;1;1;1;,792246181862098;,455172947804528;,282098992504161;,612783856719735;,342422680822206;,635483746814912;,545568635203218;,024095755024408;1,32739599775578;1,97703142341053;,0464187842760956;1,23426433472172;
 ; ; ; ; ; ; ; ; ; 
;,21;,43;,03;,23;,145512972504407;,230023084421597;,677696219362006
S.biplagiatus;2;3;3;5;3; ; ; ; ; ; ; ; 
;,836767047394205;,466125870418199;,264817823009536;,552841968657781;,260412144755717;,598164947444438;,4983105537896;,0855036278802939;1,33694283069128;2,13313765024025;,0453229787866575;
 ; ; ; ; ; ; ; ; ; ; ; ; ; ;,08; ; ; 
S.bivulneratus;2;12;2;2;3;1;0;0;0;1;0;1;0;,835043402865398;,471012851443871;,258397804095509;,537315583145303;,18820699090063;,598790506763115;,598400283482046;,0314592097715922;1,25642559919842;2,14112284487485;,0414092567968734;1,10104188722047;
 ; ; ; ; ; ; ; ; ; ;0;,31; ;,15; ; ; 
S.diffidens;2;6;1;2;1; ; ; ; ; ; ; ; 
;,608472545741434;,255573993822046;,218141468157678;,516086859963248;,143266117697832;,550439226806216;,676773523965308;,0271271526508594;1,34627046874005;2,00865053184259;,024622474908332;,964286850703799;
 ; ; ; ; ; ; ; ; ; ; ; ;,03; ; ; ; 
S.diversiventris;2;12;2;2;3;1;1;0;1;1;0;1;1;,921829483052538;,539703238947825;,322357144118318;,627365856592733;,234710235477396;,651278013998144;,544743836002248;,0326779210370329;1,2204558504511;2,11577635149136;,037265778837115;1,08711738081978;1,26951294421792;1,72181061521255;
 ; ; ; ; ; ; ; ;,085;,53;,03;,15; ; ; 
S.explanipennis;2;13;2;2;3;1;1;1;1;1;0;0;0;,712718080791599;,402652978653632;,264988255392589;,526339277389844;,24551266781415;,593286067020457;,499829714642041;,0215459014836307;1,20720417121089;1,96828755923989;,03320937723039;1,04611258064923;
 ; ; ; ; ; ; ; ; ; ;,73;,07;,09;,31; ; ; 
S.feae;2;10;3;4;3; ; ; ; ; ; ; ; 
;,81649318403725;,439937652477201;,256918094037461;,57978359661681;,264817823009536;,550228353055094;,617990952106079;,0289088250953839;1,1923269923421;2,11425780972;,0447544325135576;1,01900813819642;
 ; ; ; ; ; ; ; ; ; ; ; ;,03;,15; ; ; 
S.gestroi;2;11;3;2;3; ; ; ; ; ; ; ; 
;,902833988022287;,510394187325463;,297459573148831;,61066016308988;,230448921378274;,626340367375042;,596651136367202;,0305838403309699;1,18650548499564;2,07607958033934;,0357261061217346;1,03260305425397;
 ; ; ; ; ; ; ; ; ; ; ; ;,06;,08; ; ; 
S.humeralis;2;9;2;2;3;1;0;0;0;1;0;1;1;,843486733272765;,490239485246287;,283892413179861;,547774705387823;,271841606536499;,608526033577194;,487013161918923;,0246132330672323;1,29981727352832;1,91843170742508;,0524393704862764;1,29083595543699;1,73559889969818;2,03302144468291;
 ; ; ; ; ; ; ; ;0;,5;,06; ; ; ; 
S.liangtangi;2;18;3;4;3;0;1;1;1;1;0;0;0;,725789163577246;,406840626299901;,264266737759607;,540329474790874;,212187604403958;,568201724066995;,537551487057746;,0168809466906489;1,08069914723472;1,7279518136151;,0258967674237214;1,02804100656187;
 ; ; ; ; ; ; ; ; ; ;,17;0;,18;,15; ; ; 
S.luteomaculatus;2;3;3;5;3;0;0;1;1;1;0;0;0;,78915749191144;,475104347501863;,309630167425899;,578639209968072;,28330122870355;,595496221825574;,443797295322638;,0524451300919501;1,34688684326287;2,17454014709947;,0365956301113384;1,0572274530717;1,46089784275655;1,70926996097583;,148507737423593;,158768293549499;,452788676679383;
 ; ; ;,630563515002093; 
;,18;0;,09;,08;,00100000016666674;,0450152013563141;1,26298842592811
S.megacephalus;2;6;1;2;1; ; ; ; ; ; ; ; 
;,646344901316295;,337019752600408;,232538717530599;,536842150287963;,143014800254095;,557507201905658;,655268931344561;,0253688610375358;1,27703580940386;1,9094941336242;,0239333646690749;,930176598900266;
 ; ; ; ; ; ; ; ; ; ; ; ;,09;,15; ; ; 
S.notaculipennis;2;23;3;2;3; ; ; ; ; ; ; ; 
;,75969302045161;,434469100636792;,262315457473892;,545307116465824;,209515014542631;,593286067020457;,538593524447202;,0305453883648286;1,26374406742029;1,94102048912323;,0422705727713352;1,10475894297975;
 ; ; ; ; ; ; ; ; ; ; ; ;,09; ; ; ; 
S.pustulatus;2;19;2;3;3;1;1;0;0;1;1;1;1;,801972791064626;,456775838007211;,285962161731393;,600972895686748;,295825432703264;,636487896353365;,590661082312481;,0364683132365346;1,44013772134215;2,25246453934757;,0418946428783282;1,12814251599211;1,75281643118827;1,95568775031351;
 ; ; ; ; ; ; ; 
;,18;,81;,15;,31;,0931345839683541;,0861063637147067;,963160353136193
S.puthzianus;2;19;2;3;3; ; ; ; ; ; ; ; 
;,793678654697431;,488903085267769;,286860485522373;,680335513414563;,271841606536499;,66838591669;,594931456274357;,0487586878257387;1,33788850513889;2,0624616277024;,0392017306422158;1,03993275478569;1,50379068305718;1,80482067872116;,170830792128198;
 ;,44090773687764;,577918355727021;,201357920790331; ;1,12391676653466; ; ; ; 
;,08;,171844529083968;,652280123034403;,224890888954785
S.rorellus;2;20;2;2;3;1;1;0;0;1;1;1;1;,554541074950017;,229206303987178;,192866012766648;,48572142648158;,0969100130080564;,57287160220048;,609351055917256;,0299496036380492;1,27546254722776;1,88098813527027;,039308909711386;1,13680587931133;
 ; ; ; ; ; ; ; ; ; ;,037;,77;,27;,15; ; ; 
S.stigmatias;2;22;3;5;3; ; ; ; ; ; ; ; 
;,786431970507838;,467793029785662;,254064452914338;,553883026643874;,223892492126343;,569373909615046;,541238754331411;,0681479623342419;1,39130948953602;2,4091577890504;,0375802527675763;1,07655055492122;
 ; ; ; ; ; ; ; ; ; ; ; ;,06; ; ; ; 
S.subsimilis;2;9;2;2;3; ; ; ; ; ; ; ; 
;,768215121441203;,376394442037266;,234444967388649;,576341350205793;,315970345456918;,580057254144392;,598179742598346;,0265725097342069;1,26291141228912;1,93880830706588;,057934123883104;1,25011542590608;
 ; ; ; ; ; ; ; ; ; ; ; ; ;,08; ; ; 
S.subthoracicus;2;23;3;2;3; ; ; ; ; ; ; ; 
;,662757831681574;,375663613960885;,224014811372864;,558975090628614;,281285937468224;,636822097587174;,517739105627926;
 ; ; ;,0530101125259719;1,19139256974809; ; ; ; ; ; ; ; ; ; ; ; ; ;,08; ; ; 
S.thanonensis;2;9;2;2;3;1;1;1;0;1;0;1;1;,784467510088917;,434382585802963;,278143636692584;,553883026643874;,285719442451568;,520348220161209;,541888928845393;,0220310941440364;1,20039752285389;1,83043191526783;,0467891871339407;1,18189650439794;1,73559889969818;2,03302144468291;,075752978302897;
 ;,541386349472648; ;,848062078981481; ; ; 
;,18;,68;,18;,15;,145512972504407;,230023084421597;,677696219362006
S.virgula;2;24;3;2;3;1;1;1;1;1;0;0;1;,742432885534214;,394458685519435;,260519768326274;,568201724066995;,224677113190724;,599883072073688;,511751933013963;,0224741535532215;1,23084967658355;2,01218665595059;,0329780428669778;1,12696129150702;
 ; ; ; ; ; ; ; ; ; ;,86;,13;,12;,23; ; ; 
(((((((((((D.scabricollis:0.0281398198,D.punctiventris:0.0522260568):0.0165304832,(D.chetri:0.0324715501,D.srivichaii:0.036577528):0.0042592543):0.016112973,(D.obliquenotatus:0.0347332056,D.siwalikensis:0.0240496956):0.0356443411):0.0076514838,(D.fornicifrons:0.0673765515,D.nitidicollis:0.1004560337):0.0145728323):0.0121603904,(((D.karen:0.0504328722,D.pseudacutus:0.0471609249):2.7666e-06,D.andrewesi:0.0480975507):0.0069669242,D.tortuosus:0.0382059675):0.010811741):0.0020684997,(D.luteolunatus:0.0562242167,D.siamensis:0.1021470027):0.010950687):0.009366087,(D.betzi:0.0402231372,D.meo:0.0430754102):0.0414707113):0.0231370948,D.tubericollis:0.0883745395):0.0084178945,(((((S.amoenus:0.0018564107,S.notaculipennis:0.0010542338):0.0781434518,(S.pustulatus:0.0884092627,S.puthzianus:0.0623078578):0.0266474282):0.0198965576,((S.angusticollis:0.0131904559,S.thanonensis:0.0197882377):0.0669117612,S.humeralis:0.0679819232):0.0160558121):0.0160685651,S.rorellus:0.0968319412):0.0325738113,((((S.biplagiatus:0.0146882231,S.virgula:2.7327e-06):0.0181242578,S.luteomaculatus:0.041652629):0.0501030856,S.subthoracicus:0.0702948875):0.0192588442,((S.explanipennis:0.0669204403,S.liangtangi:0.0837475091):0.0191896571,S.stigmatias:0.0859050079):0.027375944):0.0340151206):0.0058362379):0.010800078,(((S.bivulneratus:0.0636351848,(S.diversiventris:2.4147e-06,S.gestroi:0.0641563764):0.0797494813):0.0277285246,S.subsimilis:0.0642784929):0.0167015008,S.feae:0.0607533321):0.0472946346):0.034510427,(S.diffidens:0.0021005521,S.megacephalus:0.0034191894):0.1022193644);
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