Dear Julien:
I tried the attached Rscript with mvMORPH to attain a phylogenetic
correlation matrix betwen all pairs of variables O - AQ in the
attached csv file (variables B-N are just variables that define the
assignment to various subgroups and won't be included in these
analyses). The tree is also attached.
Howver, when I am running your script under these data, I am always
getting various error messages. Maybe, I need to sort the species
lists prior to the analysis, so that both data frame and tree show the
same sequence, but the problem might be something else.
Would you mind to have a short check on my data?
Thanks a lot,
Oliver
Zitat von Julien Clavel <julien.cla...@hotmail.fr>:
Dear Oliver,
Yes, you can use the independent contrasts to estimate the
correlations (e.g., assuming BM). Otherwise, you can compute the correlations
directly from the models fit in mvMORPH.
For instance, under BM you can use:
fit_bm <- mvBM(tree, data)
cov2cor(fit_bm$sigma) # marginal correlations
cor2pcor(fit_bm$sigma) # partial correlations
Under OU, you can also retrieve the correlations from the
variance-covariance matrix
(which depends on both the “sigma” and “alpha” parameters, and can
be retrieved
with the “stationary” function):
fit_ou <- mvOU(tree, data)
cov2cor(stationary(fit_ou$sigma)) # marginal correlations
cor2pcor(stationary(fit_ou)) # partial correlations
Alternatively, you can use the “mvgls” function to do it, e.g.:
fit_bm2 <- mvgls(data~1, tree=tree, model="BM", method="LL")
cov2cor(fit_bm2$sigma$Pinv) # marginal correlations
cor2pcor(fit_bm2$sigma$Pinv) # partial correlations
You can for instance use penalized likelihood to obtain a
regularized estimate of the evolutionary correlations:
fit_bm2 <- mvgls(data~1, tree=tree, model="BM", method="PL") # Ridge
penalization by default
cov2cor(fit_bm2$sigma$Pinv) # marginal correlations
cor2pcor(fit_bm2$sigma$Pinv) # partial correlations
With LASSO penalization, for instance, you can find a sparse
estimate for the partial correlations. That is, you can directly
select the “significant”
partial correlations from the model fit:
fit_bm2 <- mvgls(data~1, tree=tree, model="BM", method="PL",
penalty="LASSO")
cov2cor(fit_bm2$sigma$Pinv) # marginal correlations
cor2pcor(fit_bm2$sigma$Pinv) # partial correlations
Best wishes,
Julien
De : R-sig-phylo <r-sig-phylo-boun...@r-project.org> de la part de
Oliver Betz <oliver.b...@uni-tuebingen.de>
Envoyé : dimanche 13 juin 2021 14:45
À : r-sig-phylo@r-project.org <r-sig-phylo@r-project.org>
Objet : [R-sig-phylo] phylogenetic correlation analysis
Dear all:
I would like to perform a phylogenetic correlation analysis (simlar to
PGLS, but correlation instead of regression), so that I get a
correlation matrix, where all the Pearson or Spearman correlation
coefficients between all of my variables are listed. One solution
might be to calculate PICs and do standard correlation analyses on
them, but there might be a more direct solution available?
Which R package would you recommend for such analysis?
Thank you very much,
Oliver Betz
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#Correlation analysis
#Data management to prepare correct data and tree file
library(mvMORPH)
setwd("c:/temp")
tree <- read.tree("DataR/transmitting science course/Own
data/Steninae_reduced.nwk")
# load eye protrusion versus attack distance towards large springtails
data <- read.csv2("DataR/transmitting science course/Own data/Daten Mittelwert
für Hauptkomponentenanalyse_Gesamtliste_Original April 2021_data
transformed_nur Relativwerte_Phyl.csv", header=T)
names(data) <- c("Species","Gattung","Artengruppe", "Untergattung",
"Klebpolstertyp",
"Tarsustyp","Hoehe_1","Hoehe_2","Hoehe_3","Hoehe_4","Litter","Rocks","Vegetation","Ground","log_Körperlänge","log_Vorderkörper","log_Pronotuml","log_rel_Hinterbeinl","log_rel_Antennenl",
"log_rel_Labiuml","LOG_eye_protrusion","log_rel_Haftpolsterfl","log_rel_Anz_Hafth_Klebp","LOG_rel_Anzahl_Haftkont_Klebp","LOG_rel_Breite_4.Vordertarsus",
"LOG_rel_Anzahl_Hafthaare_4.Vordertarsus",
"LOG_rel_Attachment_force_smooth","LOG_rel_Attachment_force_rough",
"LOG_rel_attack_dist_small_Md",
"LOG_rel_attack_dist_large_Md",
"LOG_rel_attack_dist_small_Lb", "LOG_rel_attack_dist_large_Lb",
"ARCSIN_capture_succ_Md_small", "ARCSIN_capture_succ_Md_large",
"ARCSIN_capture_succ_Lb_small",
"ARCSIN_capture_succ_Lb_large",
"Niche_width_elevation","Niche_width_habitat", "Species_freq_Doi_Inth",
"Species_freq_Doi_Php",
"ARCSIN_selection_index_vertical",
"ARCSIN_selection_index_ground", "ARCSIN_selection_index_stones") # rename
colums with shorter names, save dataframe as "dat"
rownames(data) <- data$Species # use species names to set rownames
#_________________________________________________
fit_bm <- mvBM(tree, data, model = "BM1")
cov2cor(fit_bm$sigma) # marginal correlations
cor2pcor(fit_bm$sigma) # partial correlations
Species;Gattung;Artengruppe;Untergattung;Klebpolstertyp;Tarsustyp;Hoehe_1;Hoehe_2;Hoehe_3;Hoehe_4;Litter;Rocks;Vegetation;Ground;log_Körperlänge;log_Vorderkörperl;log_Pronotuml;LOG_rel_Hinterbeinl;LOG_rel_Antennenl;LOG_rel_Labiuml;LOG_eye_protrusion;LOG_rel_Haftpolsterfl;LOG_rel_Anzahl_Hafth_Klebp;LOG_rel_Anzahl_Haftkont_Klebp;LOG_rel_Breite_4.Vordertarsus;LOG_rel_Anzahl_Hafthaare_4.Vordertarsus;LOG_rel_Attachment_force_smooth;LOG_rel_Attachment_force_rough;LOG_rel_attack_dist_small_Md;LOG_rel_attack_dist_large_Md;LOG_rel_attack_dist_small_Lb;LOG_rel_attack_dist_large_Lb;ARCSIN_capture_succ_Md_small;ARCSIN_capture_succ_Md_large;ARCSIN_capture_succ_Lb_small;ARCSIN_capture_succ_Lb_large;Niche_width_elevation;Niche_width_habitat;Species_freq_Doi_Inth;Species_freq_Doi_Php;ARCSIN_selection_index_vertical;ARCSIN_selection_index_ground;ARCSIN_selection_index_stones
D.andrewesi;1;2;
;1;2;1;1;0;0;0;1;0;0;,830588668685144;,527468782530814;,309630167425899;,642407442571537;,363878541576991;,116152348313469;,317854489331469;
; ;
;,0336209064093937;,897109673499622;1,20682587603185;1,66558099101795;,280508328519379;,424033067340483;
; ;,570199503280313;0; ; ;,17;0;,12; ; ; ;
D.betzi;1;1;
;1;1;1;1;0;1;1;1;1;1;,66254063013571;,369345755134671;,240399465738641;,554005731593748;,225172209272876;,107485819173744;,443716607769532;
; ;
;,0244340161199135;,936221491181442;2,01494034979294;2,05804623039528;,389614165156824;,295592378040602;
; ;,656060590924923;,477995198518952; ;
;,066;,11;,15;,15;,00200000133333573;,144502365941527;1,02362572892893
D.chetri;1;28;
;1;1;1;1;0;0;0;1;0;0;,870403905279027;,619441183654473;,341764159874347;,66838591669;,352316912196267;,152288344383056;,23946632295603;
; ;
;,0424724041091961;,700346372889992;1,01703333929878;1,96473092105363;,114222431987793;,452311902925352;
; ;,635276383213721;0; ; ;,37;0;,09; ; ; ;
D.fornicifrons;1;28;
;1;2;1;0;0;0;1;1;0;0;,841234295506041;,599773939146388;,328950132069784;,673020907128896;,389166084364532;,115918674280109;,217299658976496;
; ;
;,0464188947015681;,952039522330594;1,14921911265538;1,78247262416629;,157981073122323;,315952565084685;
; ;,617624310253785;,317823703927881; ;
;0;,47;,09;,08;,0140004573736748;1,40326848329556;0
D.karen;1;2;
;1;1;1;1;0;0;0;1;1;0;,704750904290671;,417969642214737;,253580289562183;,57287160220048;,330413773349191;,12057393120585;,289053557592641;
; ;
;,0364489893366717;,935587552592184;1,87448181769947;2,73663549768682;,25848569872926;,261795198153571;
; ;,20064353830785;,101273024409407; ;
;,4;,29;,27;,23;,00500002083356771;,412608195642369;,637265648700304
D.luteolunatus;1;29;
;1;2;1;1;0;0;1;1;0;0;,780965029608317;,475089803389007;,276461804173244;,571708831808688;,309630167425899;,12057393120585;,311011748981288;
; ;
;,035926420251516;,815651321782137;1,08635983067475;1,89652621748956;,198286317596186;,29190738046508;
; ;,40639442908634;,234338894401164; ;
;,47;,13;,24;,23;,00100000016666674;,210552264721877;,912441752748123
D.meo;1;1;
;1;1;1;1;0;0;0;1;0;1;,638589083292717;,353339095311305;,227886704613674;,584331224367531;,296665190261531;,0969100130080564;,445682026852679;
; ;
;,0340420429872385;,917882192744799;1,64443858946784;1,99431715266964;,228593752039125;,316486946727806;
; ;,843086960339661;,173874786487402; ;
;,43;,38;,09;,08;,00100000016666674;,205442117510134;,919007579501776
D.nitidicollis;1;28;
;1;2;1;1;1;0;1;1;0;0;,852654234153498;,610964184440695;,339593390803342;,716837723299524;,531478917042255;,136720567156407;,230266406359028;
; ;
;,0462096277662557;,697899073446361;,968482948553935;1,6222140229663;,284471504374667;,371890916572443;
; ;,430912433957024;,203705913292907; ; ;,47;,55;,15;,23; ; ;
D.obliquenotatus;1;28;
;1;2;1;1;0;0;0;1;0;1;,872331121230251;,630732892817196;,338854746252323;,684626674476734;,404503778174426;,135215190430548;,249589657772943;
; ;
;,0340062524460803;,781513275388627;1,68663626926229;2,24772783290972;,331017713245311;,366557263298232;
; ;,388715439357427;,366017949297474; ;
;,46;,17;,3;,15;,00400001066674347;1,49331728187102;0
D.pseudacutus;1;29;
;1;3;0;1;0;0;0;1;0;0;,692229835772756;,462697408101717;,283074974735472;,617000341120899;,378397900948138;,117249650556733;,27214341759105;
; ;
;,037919023916281;,914995733933636;2,02775720469055;2,58194965837332;,336191787314376;,36952675560824;
; ;,448484047723639;,338105027201183; ;
;0;0;,03;,08;,0380091512810838;,986317927412527;,128352127729054
D.punctiventris;1;28;
;1;2;1;0;0;0;0;1;0;0;,880870732532424;,574841195063385;,342027688087472;,606381365110605;,262451089730429;,126324055793894;,240649074620679;
; ; ;,0296947978482294;,759421208085003; ; ;,159589285075503;,242727209249022;
; ;,366125045404757;,237218568094435; ; ;0;,12;,09;
;,0150005625569608;1,39737400569929;0
D.scabricollis;1;28;
;1;1;1;1;0;0;1;1;0;0;,723181928261923;,455859567203536;,273001272063738;,627950765074281;,382959950139757;,147543829864141;,269785265754435;
; ;
;,0306552267004634;,83412320846046;1,22271647114758;1,76342799356294;,155249256671107;,191211403710181;
; ;,811905938993222; ; ; ;,3;,19;,09;,08; ; ;
D.siamensis;1;30; ;1;3; ; ; ; ; ; ; ;
;,649334858712142;,394451680826216;,227886704613674;,648289284739931;,423601998993174;,119054966543886;,339650157613684;
; ; ;,0639802413612563;1,04286490604421;2,20194306340165;2,13513265137677; ; ;
; ; ; ; ; ; ; ; ;,08; ; ;
D.siwalikensis;1;28;
;1;1;1;0;0;0;0;1;0;1;,87633332908638;,629205657102304;,362293937964231;,635483746814912;,338456493604605;,113943352306837;,230065551206047;
; ;
;,0299943009666998;,730857451151487;1,11727129565576;1,60852603357719;,234527383501801;,32972481522594;
; ;,609063977170518;,261055051689916; ; ;0;,17;,15; ; ; ;
D.srivichaii;1;28;
;1;2;1;1;1;0;1;1;1;0;,719993826367604;,470998169660874;,253822438708073;,661812685537261;,421603926869831;,15836249209525;,264794219407155;
; ; ;,0403563496555686;,879907418931813;1,64246452024212;2,59769518592551; ; ;
; ; ; ; ; ;,51;,091;,45;,23;,0150005625569608;1,39737400569929;0
D.tortuosus;1;31;
;1;2;1;1;0;0;0;1;0;1;,809559714635268;,525476726000246;,296299468570948;,606381365110605;,330413773349191;,12057393120585;,295786940251609;
; ;
;,0314869252496651;,824229930737481;1,40823996531185;1,84198480459011;,179872667590043;,329520882212919;
; ;,716699831265603;,131175869380332; ; ;,17;,098;,09;
;,00300000450001823;,276510115512615;,809583593564425
D.tubericollis;1;1;
;1;1;1;1;0;0;0;1;0;0;,557958104678714;,302114376956201;,222716471147583;,517195897949974;,209515014542631;,0999004309846666;,448300428150309;
; ;
;,0251009610468134;,840167747053471;1,85733249643127;2,09412159584056;,197589115496967;,339762821989302;
; ;,72856550462205;,157854757786628; ;
;,3;0;,03;,15;0;,374707134712372;,686714729593209
S.amoenus;2;9;2;2;3;1;1;0;0;1;1;1;1;,764791280484541;,43756560486588;,279223943904862;,547774705387823;,227886704613674;,599883072073688;,54139961949684;,0269985442399249;1,23711637529876;1,87528853603335;,0428105819284757;1,10703632142884;1,40654018043396;1,76715586608218;,261737849803336;,220004006354273;,434296025514549;,49162198147494;,577343568411807;,167481885486474;,793910184547671;,357357607953973;,5;,86;,12;,23;,0590342835640201;,0430132622044499;1,11520271518489
S.angusticollis;2;9;2;2;3;1;1;0;0;1;1;1;1;,792246181862098;,455172947804528;,282098992504161;,612783856719735;,342422680822206;,635483746814912;,545568635203218;,024095755024408;1,32739599775578;1,97703142341053;,0464187842760956;1,23426433472172;
; ; ; ; ; ; ; ; ;
;,21;,43;,03;,23;,145512972504407;,230023084421597;,677696219362006
S.biplagiatus;2;3;3;5;3; ; ; ; ; ; ; ;
;,836767047394205;,466125870418199;,264817823009536;,552841968657781;,260412144755717;,598164947444438;,4983105537896;,0855036278802939;1,33694283069128;2,13313765024025;,0453229787866575;
; ; ; ; ; ; ; ; ; ; ; ; ; ;,08; ; ;
S.bivulneratus;2;12;2;2;3;1;0;0;0;1;0;1;0;,835043402865398;,471012851443871;,258397804095509;,537315583145303;,18820699090063;,598790506763115;,598400283482046;,0314592097715922;1,25642559919842;2,14112284487485;,0414092567968734;1,10104188722047;
; ; ; ; ; ; ; ; ; ;0;,31; ;,15; ; ;
S.diffidens;2;6;1;2;1; ; ; ; ; ; ; ;
;,608472545741434;,255573993822046;,218141468157678;,516086859963248;,143266117697832;,550439226806216;,676773523965308;,0271271526508594;1,34627046874005;2,00865053184259;,024622474908332;,964286850703799;
; ; ; ; ; ; ; ; ; ; ; ;,03; ; ; ;
S.diversiventris;2;12;2;2;3;1;1;0;1;1;0;1;1;,921829483052538;,539703238947825;,322357144118318;,627365856592733;,234710235477396;,651278013998144;,544743836002248;,0326779210370329;1,2204558504511;2,11577635149136;,037265778837115;1,08711738081978;1,26951294421792;1,72181061521255;
; ; ; ; ; ; ; ;,085;,53;,03;,15; ; ;
S.explanipennis;2;13;2;2;3;1;1;1;1;1;0;0;0;,712718080791599;,402652978653632;,264988255392589;,526339277389844;,24551266781415;,593286067020457;,499829714642041;,0215459014836307;1,20720417121089;1,96828755923989;,03320937723039;1,04611258064923;
; ; ; ; ; ; ; ; ; ;,73;,07;,09;,31; ; ;
S.feae;2;10;3;4;3; ; ; ; ; ; ; ;
;,81649318403725;,439937652477201;,256918094037461;,57978359661681;,264817823009536;,550228353055094;,617990952106079;,0289088250953839;1,1923269923421;2,11425780972;,0447544325135576;1,01900813819642;
; ; ; ; ; ; ; ; ; ; ; ;,03;,15; ; ;
S.gestroi;2;11;3;2;3; ; ; ; ; ; ; ;
;,902833988022287;,510394187325463;,297459573148831;,61066016308988;,230448921378274;,626340367375042;,596651136367202;,0305838403309699;1,18650548499564;2,07607958033934;,0357261061217346;1,03260305425397;
; ; ; ; ; ; ; ; ; ; ; ;,06;,08; ; ;
S.humeralis;2;9;2;2;3;1;0;0;0;1;0;1;1;,843486733272765;,490239485246287;,283892413179861;,547774705387823;,271841606536499;,608526033577194;,487013161918923;,0246132330672323;1,29981727352832;1,91843170742508;,0524393704862764;1,29083595543699;1,73559889969818;2,03302144468291;
; ; ; ; ; ; ; ;0;,5;,06; ; ; ;
S.liangtangi;2;18;3;4;3;0;1;1;1;1;0;0;0;,725789163577246;,406840626299901;,264266737759607;,540329474790874;,212187604403958;,568201724066995;,537551487057746;,0168809466906489;1,08069914723472;1,7279518136151;,0258967674237214;1,02804100656187;
; ; ; ; ; ; ; ; ; ;,17;0;,18;,15; ; ;
S.luteomaculatus;2;3;3;5;3;0;0;1;1;1;0;0;0;,78915749191144;,475104347501863;,309630167425899;,578639209968072;,28330122870355;,595496221825574;,443797295322638;,0524451300919501;1,34688684326287;2,17454014709947;,0365956301113384;1,0572274530717;1,46089784275655;1,70926996097583;,148507737423593;,158768293549499;,452788676679383;
; ; ;,630563515002093;
;,18;0;,09;,08;,00100000016666674;,0450152013563141;1,26298842592811
S.megacephalus;2;6;1;2;1; ; ; ; ; ; ; ;
;,646344901316295;,337019752600408;,232538717530599;,536842150287963;,143014800254095;,557507201905658;,655268931344561;,0253688610375358;1,27703580940386;1,9094941336242;,0239333646690749;,930176598900266;
; ; ; ; ; ; ; ; ; ; ; ;,09;,15; ; ;
S.notaculipennis;2;23;3;2;3; ; ; ; ; ; ; ;
;,75969302045161;,434469100636792;,262315457473892;,545307116465824;,209515014542631;,593286067020457;,538593524447202;,0305453883648286;1,26374406742029;1,94102048912323;,0422705727713352;1,10475894297975;
; ; ; ; ; ; ; ; ; ; ; ;,09; ; ; ;
S.pustulatus;2;19;2;3;3;1;1;0;0;1;1;1;1;,801972791064626;,456775838007211;,285962161731393;,600972895686748;,295825432703264;,636487896353365;,590661082312481;,0364683132365346;1,44013772134215;2,25246453934757;,0418946428783282;1,12814251599211;1,75281643118827;1,95568775031351;
; ; ; ; ; ; ;
;,18;,81;,15;,31;,0931345839683541;,0861063637147067;,963160353136193
S.puthzianus;2;19;2;3;3; ; ; ; ; ; ; ;
;,793678654697431;,488903085267769;,286860485522373;,680335513414563;,271841606536499;,66838591669;,594931456274357;,0487586878257387;1,33788850513889;2,0624616277024;,0392017306422158;1,03993275478569;1,50379068305718;1,80482067872116;,170830792128198;
;,44090773687764;,577918355727021;,201357920790331; ;1,12391676653466; ; ; ;
;,08;,171844529083968;,652280123034403;,224890888954785
S.rorellus;2;20;2;2;3;1;1;0;0;1;1;1;1;,554541074950017;,229206303987178;,192866012766648;,48572142648158;,0969100130080564;,57287160220048;,609351055917256;,0299496036380492;1,27546254722776;1,88098813527027;,039308909711386;1,13680587931133;
; ; ; ; ; ; ; ; ; ;,037;,77;,27;,15; ; ;
S.stigmatias;2;22;3;5;3; ; ; ; ; ; ; ;
;,786431970507838;,467793029785662;,254064452914338;,553883026643874;,223892492126343;,569373909615046;,541238754331411;,0681479623342419;1,39130948953602;2,4091577890504;,0375802527675763;1,07655055492122;
; ; ; ; ; ; ; ; ; ; ; ;,06; ; ; ;
S.subsimilis;2;9;2;2;3; ; ; ; ; ; ; ;
;,768215121441203;,376394442037266;,234444967388649;,576341350205793;,315970345456918;,580057254144392;,598179742598346;,0265725097342069;1,26291141228912;1,93880830706588;,057934123883104;1,25011542590608;
; ; ; ; ; ; ; ; ; ; ; ; ;,08; ; ;
S.subthoracicus;2;23;3;2;3; ; ; ; ; ; ; ;
;,662757831681574;,375663613960885;,224014811372864;,558975090628614;,281285937468224;,636822097587174;,517739105627926;
; ; ;,0530101125259719;1,19139256974809; ; ; ; ; ; ; ; ; ; ; ; ; ;,08; ; ;
S.thanonensis;2;9;2;2;3;1;1;1;0;1;0;1;1;,784467510088917;,434382585802963;,278143636692584;,553883026643874;,285719442451568;,520348220161209;,541888928845393;,0220310941440364;1,20039752285389;1,83043191526783;,0467891871339407;1,18189650439794;1,73559889969818;2,03302144468291;,075752978302897;
;,541386349472648; ;,848062078981481; ; ;
;,18;,68;,18;,15;,145512972504407;,230023084421597;,677696219362006
S.virgula;2;24;3;2;3;1;1;1;1;1;0;0;1;,742432885534214;,394458685519435;,260519768326274;,568201724066995;,224677113190724;,599883072073688;,511751933013963;,0224741535532215;1,23084967658355;2,01218665595059;,0329780428669778;1,12696129150702;
; ; ; ; ; ; ; ; ; ;,86;,13;,12;,23; ; ;
(((((((((((D.scabricollis:0.0281398198,D.punctiventris:0.0522260568):0.0165304832,(D.chetri:0.0324715501,D.srivichaii:0.036577528):0.0042592543):0.016112973,(D.obliquenotatus:0.0347332056,D.siwalikensis:0.0240496956):0.0356443411):0.0076514838,(D.fornicifrons:0.0673765515,D.nitidicollis:0.1004560337):0.0145728323):0.0121603904,(((D.karen:0.0504328722,D.pseudacutus:0.0471609249):2.7666e-06,D.andrewesi:0.0480975507):0.0069669242,D.tortuosus:0.0382059675):0.010811741):0.0020684997,(D.luteolunatus:0.0562242167,D.siamensis:0.1021470027):0.010950687):0.009366087,(D.betzi:0.0402231372,D.meo:0.0430754102):0.0414707113):0.0231370948,D.tubericollis:0.0883745395):0.0084178945,(((((S.amoenus:0.0018564107,S.notaculipennis:0.0010542338):0.0781434518,(S.pustulatus:0.0884092627,S.puthzianus:0.0623078578):0.0266474282):0.0198965576,((S.angusticollis:0.0131904559,S.thanonensis:0.0197882377):0.0669117612,S.humeralis:0.0679819232):0.0160558121):0.0160685651,S.rorellus:0.0968319412):0.0325738113,((((S.biplagiatus:0.0146882231,S.virgula:2.7327e-06):0.0181242578,S.luteomaculatus:0.041652629):0.0501030856,S.subthoracicus:0.0702948875):0.0192588442,((S.explanipennis:0.0669204403,S.liangtangi:0.0837475091):0.0191896571,S.stigmatias:0.0859050079):0.027375944):0.0340151206):0.0058362379):0.010800078,(((S.bivulneratus:0.0636351848,(S.diversiventris:2.4147e-06,S.gestroi:0.0641563764):0.0797494813):0.0277285246,S.subsimilis:0.0642784929):0.0167015008,S.feae:0.0607533321):0.0472946346):0.034510427,(S.diffidens:0.0021005521,S.megacephalus:0.0034191894):0.1022193644);
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