On Thu, May 28, 2015 at 05:07:53PM -0700, Dario Copetti wrote:
> Getting to the practical point, I took the value in col 9 of the
> reads that have either a flag = 83 or 99, and the distribution is
> similar to what I expect (between 30 and 40 kb). If I take the col 9
> values of the reads with flag 97 or 81, 99% of the hits are either
> below 5 kb or above 90 kb. aren't these still pairs of reads,
> aligned towards each other?

The proper-pair flag (2) is set by the aligner and it is up to that
software to describe precisely how it defines a proper pair. 

Your own analysis seems to imply it is using the insert size as a tool
for distinguishing between proper pair and not.  Some aligners will
subsample; so the first N reads get aligned and from this the insert
size distribution is determined and normal vs abnormal insert sizes
are then chosen.[1]

However it really is outside the scope of SAM spec itself, so the
aligner specific help forums would be better served to your query.

James

[1] This is why it is recommended that any realignment task - taking an
existing set of aligned data and realigning using new parameters - is
done with data that is not in a genome specific order (eg use name
sorted instead) to avoid any genomic location biases on insert size.

-- 
James Bonfield ([email protected]) | Hora aderat briligi. Nunc et Slythia Tova
                                  | Plurima gyrabant gymbolitare vabo;
  A Staden Package developer:     | Et Borogovorum mimzebant undique formae,
https://sf.net/projects/staden/   | Momiferique omnes exgrabure Rathi. 


-- 
 The Wellcome Trust Sanger Institute is operated by Genome Research 
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