As far as I can see, envelope-from domains without a SOA record see their email 
rejected with 554 over some volume threshold.
This seems to be a fairly recent update. This requirement only recently 
appeared here: https://senders.yahooinc.com/smtp-error-codes/ (you can check 
the internet archive for the previous version which didn’t mention SOA).

Rodolfo Saccani
CTO @ Libraesva


Da: mailop <mailop-boun...@mailop.org> per conto di Jarland Donnell via mailop 
<mailop@mailop.org>
Data: mercoledì, 19 aprile 2023, 04:43
A: mailop@mailop.org <mailop@mailop.org>
Oggetto: Re: [mailop] [E] Yahoo "Message not allowed" error
That is actually the entirety of the SMTP response from
mx-eu.mail.am0.yahoodns.net at that time:

"554 Message not allowed - [299]"

Other error messages linked to
 
https://urlsand.esvalabs.com/?u=https%3A%2F%2Fpostmaster.yahooinc.com%2Ferror-codes&e=20266bc5&h=cb659c4b&f=y&p=y
  but not that one. Nothing
seems to help much on that page for this case unless I just assume the
message is "Those people can't email Yahoo anymore." I'm fairly
confused.

On 2023-04-18 16:25, Marcel Becker via mailop wrote:
> On Tue, Apr 18, 2023 at 12:16 PM Jarland Donnell via mailop
> <mailop@mailop.org> wrote:
>
>> id=<481770.862217189-sendEmail@srv4414> (554 Message not allowed -
>> [299])
>
> This is most likely not the full SMTP response we provide. The full
> one will tell you the reason -- including a link to a page explaining
> what to do and who to contact.
>
> -- Marcel
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> mailop mailing list
> mailop@mailop.org
>  
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