Hi Listers,

I am trying to better understand how spatial autocorrelation is determined.
I have attached 2 txt files one is a fish species x sites abundance table
and the other is a distance file with the lat and longs for each site
(converted to cartesian coords). I have run the tests for spatial
correlation using the methodology from the Borcard et al. Use R! book.

The result was no significant spatial correlation. However an nmds
ordination of the fish data shows 3 very strongly clustered groups of sites
which corresponds to 3 oceanic islands. A permutation analysis of the fish
resemblance matrix with a distance matrix was also highly significant with
a rho=0.646 spearmans rank correlation. To my mind I would have thought
that with such strong groupings in the data that spatial correlation would
be present at the scale of islands? Can someone help me explain this result?

My analysis below

fish_hellinger <- decostand (fish,"hellinger")
fish.xy # spatial dataset space_ed.txt

#test for overall trend
lineartest <- anova(rda(fish_hellinger,fish.xy))

Permutation test for rda under reduced model

Model: rda(X = fish_hellinger, Y = fish.xy)
         Df     Var      F N.Perm Pr(>F)
Model     2 0.17148 8.6098    199  0.005 ** ##significant linear trend
Residual 14 0.13942
---
Signif. codes:  0 ‘***’ 0.001 ‘**’ 0.01 ‘*’ 0.05 ‘.’ 0.1 ‘ ’ 1

 # fit linear trend to the fish data and then extract residuals for
subsequent use in spatial analysis
> fish_hellinger_detrended <- resid(lm(as.matrix(fish_hellinger)~
.,data=fish.xy))

# make a distance matrix from the residuals data for input to the
correlogram call
> fish_hellinger_D1 <- dist(fish_hellinger_detrended)

# perform the mantel-based test for spatial correlation
> (fish_correlogram <-
mantel.correlog(fish_hellinger_D1,XY=fish.xy,nperm=99))

Mantel Correlogram Analysis

Call:

mantel.correlog(D.eco = fish_hellinger_D1, XY = fish.xy, nperm = 99)

        class.index      n.dist  Mantel.cor Pr(Mantel) Pr(corrected)
D.cl.1   118.013676  462.000000   -0.042424       0.11          0.11
D.cl.2   352.037024    0.000000          NA         NA            NA
D.cl.3   586.060371    0.000000          NA         NA            NA
D.cl.4   820.083719    0.000000          NA         NA            NA
D.cl.5  1054.107067  384.000000    0.033639       0.30            NA
D.cl.6  1288.130414    0.000000          NA         NA            NA
D.cl.7  1522.153762    0.000000          NA         NA            NA
D.cl.8  1756.177109  320.000000          NA         NA            NA
D.cl.9  1990.200457    0.000000          NA         NA            NA
D.cl.10 2224.223804    0.000000          NA         NA            NA
D.cl.11 2458.247152  240.000000          NA         NA            NA


-- 
cheers

Andy


Andrew Halford Ph.D
Research Scientist (Kimberley Marine Parks)
Dept. Parks and Wildlife
Western Australia

Ph: +61 8 9219 9795
Mobile: +61 (0) 468 419 473
LUT_GIBB        LUT_FULV        CHL_STRO        SCA_PRAS        NAS_LITU        
KYP_VAIG        LUT_BOHA        SCA_RUBR        NAS_UNIC        HIP_HARI        
APH_FURC        NAS_ELEG        CHL_ENNE        KYP_CINE        NAS_BREV        
CEP_ARGU        BOL_MURI        LUT_KASM        CHL_MICR        LET_OBSE        
LUT_DECU        VAR_LOUT        CET_BICO        LUT_MONO        NAS_VLAM        
HIP_LONG        CHE_UNDU        GRA_ALBO        NAS_CAES        ACA_XANT        
LUT_RIVU        CEP_MINI        COR_AYGU        COR_GAIM        LET_XANT        
TRI_OBES        NAS_FAGE        PMS_LAEV        LET_ERUS        LET_OLIV        
CAC_MELA        CAC_AMBL        EPI_FUSC        NAS_HEXA        PMS_AREO        
APR_VIRE        SCA_GHOB        SCA_XANT        EPI_TAUV        EPI_POLY        
EPI_MACR        LET_EHUS        AET_ROGA        NAS_TONG        VAR_ALBI        
LET_HARA        SYM_NEMA        
Coc1    1       6       16      5       9       0       6       1       20      
17      0       9       0       14      3       0       0       0       0       
10      0       1       0       3       0       0       0       0       1       
0       0       0       0       0       0       0       0       0       1       
0       1       0       1       0       0       0       1       0       0       
0       0       0       0       0       0       0       0
Coc4    0       0       1       24      12      0       0       7       4       
24      0       15      0       0       0       0       0       0       0       
0       0       0       0       0       0       0       0       0       0       
0       0       0       0       0       0       1       0       0       0       
1       2       1       0       0       0       1       1       0       0       
0       0       0       0       0       0       0       0
Coc8    5       21      22      19      15      6       4       6       8       
11      0       28      0       3       7       0       0       0       0       
5       0       1       0       2       0       0       2       0       0       
0       0       0       2       2       0       0       0       0       0       
0       1       0       0       0       0       0       0       0       0       
0       2       0       0       0       0       0       0
Coc12   0       0       40      20      6       0       0       5       6       
19      0       16      4       0       0       0       0       0       0       
2       0       3       0       0       0       0       0       0       0       
0       0       0       0       0       1       0       0       0       0       
0       1       0       0       0       0       0       0       0       0       
0       1       0       0       0       0       0       0
Coc13   13      43      31      2       7       59      4       7       8       
44      0       9       43      49      0       0       0       0       0       
0       0       0       0       0       0       0       3       0       0       
6       0       0       2       0       0       1       0       0       0       
0       2       0       1       0       0       0       1       0       0       
0       0       0       0       0       0       0       0
Coc16   96      89      34      30      4       4       0       3       3       
20      0       5       8       5       0       0       0       0       0       
6       0       0       0       0       0       0       1       0       0       
0       0       0       0       0       1       0       0       0       0       
0       0       1       0       0       0       0       1       0       0       
0       0       0       0       0       0       4       0
Coc18   3       3       20      26      13      0       1       0       0       
15      0       8       0       0       0       1       0       0       0       
0       0       1       0       3       8       0       1       1       0       
0       0       0       1       0       0       2       0       0       0       
0       0       5       3       0       0       0       1       0       0       
0       1       0       0       0       0       0       0
Coc23   45      10      47      210     8       18      5       4       1       
14      0       9       94      14      0       0       1       0       0       
0       0       0       0       6       0       0       1       0       0       
0       0       0       2       0       1       5       0       0       0       
0       2       1       0       0       0       0       0       0       1       
0       2       0       0       0       0       0       0
Coc25   0       186     22      15      5       0       0       5       8       
16      0       3       11      0       0       0       2       0       0       
0       0       2       0       0       0       0       5       0       0       
0       0       0       0       0       1       0       0       0       0       
0       2       0       0       0       0       0       1       0       0       
0       0       0       0       0       0       0       0
Coc28   44      452     78      30      10      0       1       4       9       
20      0       16      12      0       0       0       0       0       0       
0       0       2       0       2       0       0       2       0       0       
0       0       0       0       0       5       0       0       0       0       
0       2       0       2       0       0       0       0       0       0       
0       0       0       0       0       0       0       0
Coc34   5       13      21      17      6       0       7       2       5       
29      0       15      10      0       0       1       0       0       0       
14      0       0       0       0       0       0       0       0       0       
0       0       0       3       0       0       0       0       0       1       
0       1       0       0       0       0       0       1       0       0       
0       1       0       0       0       0       0       0
Coc35   0       2       17      21      12      0       3       3       38      
9       0       17      0       0       14      2       2       2       0       
8       0       1       0       0       2       0       1       1       1       
0       0       0       0       0       1       0       0       0       0       
0       2       2       2       0       0       0       1       0       0       
0       0       0       0       0       0       0       0
X1      0       0       1       0       0       38      13      11      0       
0       14      0       1       17      6       4       0       0       0       
0       0       1       0       0       3       0       0       12      15      
0       0       5       0       0       0       0       0       0       0       
0       0       1       0       0       0       0       0       1       0       
0       0       0       0       0       0       0       0
X2      0       0       0       0       7       12      11      12      0       
0       13      0       0       6       0       5       0       0       0       
0       0       3       0       0       0       0       0       0       3       
0       1       0       0       0       0       0       0       0       0       
0       0       0       0       0       0       0       0       0       0       
0       0       0       0       0       0       0       0
X4      0       0       0       0       8       16      3       16      1       
0       22      1       0       1       0       5       0       13      0       
0       0       3       0       0       0       0       0       2       0       
1       0       10      0       3       0       0       0       0       0       
0       0       0       0       1       0       0       0       0       0       
0       0       0       0       0       0       0       0
X7      0       0       2       0       2       0       5       21      0       
0       21      0       0       0       8       11      0       0       2       
0       0       6       0       0       0       0       0       0       0       
0       1       0       1       2       0       0       0       0       0       
0       0       0       0       2       0       0       0       1       0       
0       0       0       0       0       0       0       0
X9      0       0       8       0       7       1       3       29      0       
0       29      0       5       0       1       5       0       0       0       
0       0       9       0       0       0       0       0       0       26      
0       0       0       1       0       1       0       0       0       0       
0       0       0       0       0       0       0       0       0       0       
0       0       0       0       0       0       0       0
X10     2       0       3       3       5       6       10      15      0       
0       20      0       0       0       2       10      0       0       0       
0       0       5       1       0       3       0       1       4       3       
0       0       5       0       0       0       0       0       0       0       
0       0       0       0       2       0       0       0       0       0       
0       0       0       0       0       1       0       0
X11     1       0       7       3       10      8       17      13      0       
0       24      0       0       2       2       0       0       0       0       
0       0       4       0       0       0       0       0       10      5       
0       0       10      0       4       0       0       0       0       0       
0       0       0       0       1       0       0       0       6       0       
0       0       0       0       0       0       0       0
X12     0       0       2       0       9       0       6       16      0       
0       21      0       0       0       0       5       0       0       0       
0       0       5       1       0       0       0       0       5       2       
0       0       6       2       0       0       0       0       0       0       
0       0       0       0       6       0       0       0       0       0       
0       0       0       0       0       1       0       0
X14     0       0       0       1       19      36      4       101     4       
0       20      0       0       0       2       1       0       0       0       
0       0       3       0       0       0       0       0       0       0       
2       0       0       0       0       0       0       0       0       0       
0       0       0       0       0       0       0       0       2       0       
0       0       0       0       4       0       0       0
X16     0       0       0       0       8       16      8       16      2       
0       26      0       0       8       8       4       0       0       0       
0       0       3       0       0       2       0       0       5       4       
0       0       5       0       0       0       0       0       0       0       
0       0       0       0       1       0       0       0       0       0       
0       0       0       0       0       0       0       0
X18     0       0       4       0       8       5       10      31      4       
0       45      0       0       0       2       0       0       0       0       
0       0       1       0       0       0       0       0       0       2       
0       0       0       0       1       0       0       0       0       0       
0       0       0       0       0       0       1       0       3       0       
0       0       0       0       0       0       0       0
X19     1       0       0       0       7       3       9       29      0       
0       10      0       0       0       1       3       0       0       0       
0       0       7       0       1       0       0       0       0       1       
0       0       0       0       2       1       1       0       0       0       
0       0       0       0       0       0       0       0       0       0       
0       0       0       0       1       2       0       0
X20     0       0       0       0       11      52      18      29      2       
0       19      0       0       0       4       0       0       2       0       
0       0       7       0       0       0       0       0       2       0       
0       0       0       0       3       0       0       0       0       0       
0       0       0       0       0       0       0       0       0       0       
0       0       0       1       0       0       0       0
X21     16      0       0       0       14      40      14      15      1       
0       21      2       0       2       3       6       0       1       0       
0       0       14      0       0       0       0       0       3       2       
0       0       0       0       2       0       1       0       0       0       
0       0       2       0       0       0       0       0       1       0       
0       0       0       0       0       0       0       0
X22     0       0       0       0       14      48      18      17      1       
0       30      0       0       0       0       7       0       12      0       
0       0       2       0       0       0       0       0       0       0       
2       0       3       0       6       0       0       0       0       0       
0       0       0       0       0       0       0       0       0       0       
0       0       0       0       0       0       0       0
X24     0       0       2       5       6       12      8       7       0       
0       26      0       0       2       0       7       0       0       0       
0       0       5       0       0       0       0       0       4       2       
0       0       0       0       1       0       0       0       0       0       
0       0       0       0       1       0       2       0       0       0       
0       0       0       1       0       0       0       0
Cl1     11      0       0       4       9       0       22      0       21      
0       3       0       0       0       4       11      12      23      10      
0       8       0       10      0       15      9       1       0       0       
41      0       0       1       0       4       0       13      6       3       
1       0       1       0       8       1       3       0       0       0       
0       0       0       0       0       0       0       0
Cl2     2       0       0       2       19      0       6       0       7       
0       3       0       0       0       0       12      10      70      4       
1       8       1       7       0       6       2       4       2       0       
0       0       0       1       2       1       1       1       1       4       
0       0       0       0       0       0       0       0       0       0       
0       0       0       1       0       0       0       0
Cl3     3       0       0       0       21      0       1       0       1       
0       7       0       0       0       41      4       0       0       6       
0       9       2       25      0       3       0       2       0       0       
0       0       0       1       2       2       0       0       0       8       
4       0       0       0       0       0       1       0       0       0       
0       0       0       0       0       0       0       0
Cl5     83      0       0       1       19      0       18      5       10      
0       7       0       0       0       19      15      0       1       29      
0       41      0       7       0       0       9       6       0       0       
0       0       0       0       2       1       1       0       5       0       
1       0       0       0       0       1       0       0       0       0       
0       0       1       0       1       0       0       0
Cl6     70      0       0       1       46      0       46      4       0       
0       4       0       0       0       12      7       0       0       12      
0       21      1       12      0       6       7       9       0       0       
0       4       0       2       1       1       3       2       2       2       
1       0       2       0       1       1       0       0       1       0       
0       0       0       0       0       0       0       2
Imp1    61      0       0       0       2       3       13      0       15      
0       4       0       0       0       5       12      7       7       6       
1       13      3       10      0       3       2       5       0       0       
18      6       0       5       1       0       1       0       1       0       
3       0       1       0       0       0       2       0       0       0       
0       0       0       0       0       0       0       0
Imp2    9       0       0       2       18      2       9       4       1       
0       6       0       0       1       0       2       1       0       31      
0       13      0       3       1       0       18      6       0       0       
0       1       0       2       0       2       0       0       3       0       
2       0       0       1       0       2       3       0       0       0       
0       0       0       1       0       0       0       0
Imp4    6       0       0       2       22      0       9       0       20      
0       7       0       0       0       9       12      0       0       24      
0       13      4       16      0       0       0       3       0       0       
0       0       0       2       3       0       0       0       0       1       
6       0       0       0       0       6       0       0       0       0       
5       0       0       0       0       0       0       0
Imp7    15      0       0       0       6       0       11      1       47      
0       4       0       0       4       13      10      0       0       16      
0       15      0       10      0       6       2       5       0       0       
3       14      0       0       0       2       2       0       7       1       
2       0       2       0       1       0       1       0       0       2       
0       0       5       1       0       0       0       0
Imp9    195     0       0       0       16      6       13      6       32      
0       1       0       0       6       16      10      11      0       5       
0       5       2       11      0       0       7       4       0       0       
3       0       0       2       0       0       3       8       3       0       
3       0       3       0       0       1       1       0       0       0       
0       0       0       0       0       0       0       0
LAT     LONG    
Coc1    10780.76        -1335.326
Coc4    10779.5 -1335.06
Coc8    10785.59        -1337.317
Coc12   10780.08        -1336.277
Coc13   10777.29        -1342.804
Coc16   10777.62        -1345.016
Coc18   10778.98        -1339.375
Coc23   10780.52        -1337.295
Coc25   10778.78        -1348.324
Coc28   10780.47        -1350.227
Coc34   10784.83        -1336.664
Coc35   10785.78        -1338.755
X1      11750.95        -1155.076
X2      11754.07        -1156.99
X4      11750.57        -1158.306
X7      11747.88        -1162.078
X9      11749.43        -1155.939
X10     11763.62        -1151.669
X11     11764.79        -1151.835
X12     11766.23        -1151.47
X14     11767.03        -1153.384
X16     11767.1 -1155.994
X18     11766.98        -1158.239
X19     11765.28        -1159.777
X20     11763.99        -1162.885
X21     11762.99        -1166.181
X22     11759.46        -1156.282
X24     11762.2 -1153.782
Cl1     13287.19        -1908.418
Cl2     13286.38        -1907.632
Cl3     13288.95        -1911.594
Cl5     13283.59        -1909.513
Cl6     13282.85        -1911.118
Imp1    13241.55        -1935.344
Imp2    13242.79        -1935.809
Imp4    13243.45        -1937.547
Imp7    13237.1 -1941.398
Imp9    13235.16        -1944.386
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