The release notes for SM 2.53.3 say:

 SeaMonkey 2.53.3 shares most parts of the mail and news code with Thunderbird. 
Please read the Thunderbird 60.0 release notes for specific changes and 
security fixes in this release.

The Thunderbird 60.0 release notes say:
Native notifications on Linux are now re-enabled

But not in SM 2.53.3, apparently (LXLE Ubuntu 16.04 x86), where I still have to use the GNotifier extension to get anything other than a sound.

/df

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