I also had this issue because firefox had grabbed my sound device
directly, and other processes couldn't access it. Rhythmbox would appear
to play ok (debug output didn't think there were any problems), however
the track time would stay at 0:00 and nothing would play.

Firefox has sound device:

$ sudo fuser -v /dev/dsp* /dev/snd/* /dev/seq*
                     USER PID ACCESS COMMAND
/dev/snd/controlC0:  <myusername>   2307 F.... pulseaudio
/dev/snd/controlC1:  <myusername>   2307 F.... pulseaudio
/dev/snd/pcmC0D0p:   <myusername>   3074 F...m firefox

I used gstreamer-properties to change the default audio output from
"Autodetect" to "PulseAudio Sound Server", restarted firefox, and
afterwards, firefox was using pulseaudio:

$ sudo fuser -v /dev/dsp* /dev/snd/* /dev/seq*
                     USER PID ACCESS COMMAND
/dev/snd/controlC0:  <myusername>   2307 F.... pulseaudio
/dev/snd/controlC1:  <myusername>   2307 F.... pulseaudio
/dev/snd/pcmC0D0p:   <myusername>   2307 F...m pulseaudio

Rhythmbox is also now able to play audio as the same time as firefox.

$ lsb_release -a
No LSB modules are available.
Distributor ID: Ubuntu
Description:    Ubuntu 9.10
Release:        9.10
Codename:       karmic

$ apt-cache policy firefox
firefox:
  Installed: 3.5.7+nobinonly-0ubuntu0.9.10.1
  Candidate: 3.5.7+nobinonly-0ubuntu0.9.10.1
  Version table:
 *** 3.5.7+nobinonly-0ubuntu0.9.10.1 0
        500 http://gb.archive.ubuntu.com karmic-updates/main Packages
        500 http://security.ubuntu.com karmic-security/main Packages
        100 /var/lib/dpkg/status
     3.5.3+build1+nobinonly-0ubuntu6 0
        500 http://gb.archive.ubuntu.com karmic/main Packages

$ apt-cache policy pulseaudio
pulseaudio:
  Installed: 1:0.9.19-0ubuntu4.1
  Candidate: 1:0.9.19-0ubuntu4.1
  Version table:
 *** 1:0.9.19-0ubuntu4.1 0
        500 http://gb.archive.ubuntu.com karmic-updates/main Packages
        100 /var/lib/dpkg/status
     1:0.9.19-0ubuntu4 0
        500 http://gb.archive.ubuntu.com karmic/main Packages

-- 
[Koala] problem with sound in totem, rhytmbox, but no in firefoxes flash videos
https://bugs.launchpad.net/bugs/405405
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