I seem to be affected by this problem also.
When it happens i see either of two error messages in dmesg: 
ath5k phy0: can't alloc skbuff of size 2559 
and
ath5k phy0: noise floor calibration timeout .....

The first error message lead me to believe that there might be an issue with 
running out of memory. 
Running top when downloading a large file over WLAN i can see that the memory 
is reducing rapidly. 
Then when the memory is almost out top freezes for a while and all of a sudden 
there is a large chunk of memory available again, that time the download didn't 
fail but i beleive that if the timing is off then it might.

So my assumption is that a high speed wlan download, the problem is more
severe when loading large chunks of data from local server or streaming
video from local server, combined with a slow disk write speed and small
RAM is the problem here.

-- 
Connection droped or system crash whean trying to transfer large files over 
wi-fi
https://bugs.launchpad.net/bugs/389891
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