For me; 
A fix on the non-performing machine mentioned on he previous post.
 Take the .wav file that I had chosen (which did not play) and 
Using  "OggConvert" convert it into an .ogg file.
OggConvert is a GUI application from the repository that 's pretty well 
intuitive.(you might have to install it)
 Go to thunderbird /edit/preferences/general  and choose  the newly created 
.ogg file from where you saved it.
Worked for me. Although I don't know which player did the job.
Now chosen .OGG sound file plays.
Just tried it on a whim and success,,,,,,,,,,,, if it lasts.

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https://bugs.launchpad.net/bugs/292676

Title:
  Thunderbird doesn't play any sound on notification

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