It is my experience also, that inlining gives the greatest performance
gain for functions that expect primitive arguments.
As Chouser said, doing this eliminates the boxing/unboxing overhead.

Here's my take on this:
The Java method signatures created by Clojure will always be Objects
in order to maintain a consistent interface. Therefore it doesn't make
sense for HotSpot to remove that boxing overhead. HotSpot sees a
"function that takes a Integer". Why should it eliminate the boxing
and treat it like a "function that takes a int"? What if the function
really did want a Integer instead of an int? This isn't a decision the
compiler can make.

Hope that helps (and is correct).
-Patrick
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