Take a look here
http://code.google.com/appengine/docs/java/config/webxml.html#Security_and_Authentication

And then here (the first link will also point you here)
http://code.google.com/appengine/docs/java/users/

Also read the overview 
http://code.google.com/appengine/docs/java/users/overview.html

The overview explains how users are authenticated.  Once a user is 
authenticated, UserService will know about the user.

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