Dear All
We are pleased to share our new open-access publication, where we use 
high-resolution dive data from beaked whales (Ziphius cavirostris) to model 
blood and tissue O₂ and CO₂ dynamics.

Beyond advancing our understanding of diving physiology, this work provides a 
mechanistic link between individual metabolic limits and broader conservation 
frameworks such as Population Consequences of Disturbance (PCoD). Our results 
show that Ziphius often exceed their calculated aerobic limits and rely on 
anaerobic metabolism, meaning that even subtle disturbances, such as sonar 
exposure or altered recovery patterns, can push animals beyond physiological 
thresholds, with potential fitness costs that accumulate at the population 
level. By integrating physiology with behavioral and ecological models, this 
approach offers a more powerful tool for predicting the impacts of disturbance 
and cumulative stressors on deep-diving cetaceans.

The abstract and a link to the article are provided below. If you experience 
any difficulty accessing the paper, please do not hesitate to contact me at 
[email protected].

Title: Modelling the effect of varying metabolic rate and cardiac output on 
estimated tissue and bloodO2 and CO2 levels in an extreme deep-diver, the 
goose-beaked whale (Ziphius cavirostris)

Authors: Andreas Fahlman, A. Schorr, G.S., Sweeney, D.A., Rone, B.K., Coates, 
S.N., Allen, A.S., Martín López, L.M., Jarvis, S.M., Falcone, E.A.

Journal: Experimental Physiology

doi: http://doi.org/10.1113/EP093021

Abstract: High-resolution movement data from Cuvier’s beaked, or goose-beaked 
whale (Ziphius cavirostris, hereafter Ziphius, n = 8) tag deployments (4.1–19.2 
days) were used to estimate blood and tissue O2 and CO2 levels. Acceleration 
and magnetometry data were used to estimate the locomotion cost (LC) from the 
relationship between activity and the O2 consumption rate. We estimated that 
the diving metabolic rate (DMR) decreased with increasing dive duration, 
ranging from 6.18 mL O2 min−1 kg−1 for very short dives (<1.0 min) to 1.65 mL 
O2 min−1 kg−1 and 2.06 mL O2 min−1 kg−1 for intermediate (>17.5 and ≤33.3 min) 
and long dives (>33.3min), respectively. The calculated aerobic dive limit 
(cADL), average behavioural ADL (bADL) and dynamic ADL (dADL) were 62.4, 61.3 
(44.3–75.4) and 41.7 (2.0–102.5) min, respectively. Despite the physiological 
and metabolic adjustments assumed by the model, the muscle O2 ran out for many 
of the stereotypical long, deep dives exhibited by these animals. Based on the 
model results, we speculate that a large portion of the foraging dives in 
Ziphius are fuelled by alternative metabolic pathways, for example, 
phosphocreatine or glycolysis. A reliance on these alternative metabolic 
pathways during foraging may require long recovery periods, including primarily 
aerobic dives. Disturbing this normal dive pattern may disrupt this normal dive 
pattern, leading to behavioural and physiological changes that could cause 
trauma.
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