Reading here (sorry, not on the 'live' site yet, due to a mistake our end):
http://test.plants.ensembl.org/Hordeum_vulgare/Info/Annotation/#assembly

~2.6 million sequenced contigs were generated using whole-genome
shotgun sequencing (WGS). ~138,000 of these are assigned to specific
chromosomal positions. These are shown in the browser on chromosomes
labeled 1-7. An additional 355,000 WGS contigs could be assigned to a
specific chromosome arm, but not to a more specific location. These
are shown in the browser on chromosomes labeled with an "H." (e.g. 1H,
3HL). "HL" and "HS" refer to chromosome long and short arms
respectively.

I was mentioning the index issue (537 Mbp), which as I'm sure you know
is fixed with the alternative index format. However, fixed or not, I
found the comments amusing :-)


Cheers,
Dan.



On 23 July 2014 11:48, John Marshall <[email protected]> wrote:
> On 23 Jul 2014, at 10:58, Dan Bolser <[email protected]> wrote:
>>
>> I like these comments:
> [...]
>>
>>        "Hopefully 2 Gbp [per chromosome - JM] suffices for most species that 
>> people are
>> interested in." - John Marshall.  [Somewhat taken out of context, see below 
>> - JM]
>>
>> As already mentioned, bread wheat, which provides ~20% of all the
>> calories consumed by humans globally, has 6 of it's 27 chromosomes
>> *assembled* to a length of greater than 537 Mbp [1]...
>
> I guess your point is that humans are therefore interested in bread wheat, 
> but I would note from your spreadsheet that 2B's 696 Mbp is substantially 
> less than 2 Gbp -- so I'm not sure what your point is after all.
>
>> I won't mention Paris japonica ;-)
>
> If I'm reading wikipedia correctly, 150 billion bp in 40 chromosomes implies 
> at least one chromosome of at least 3.75 Gbp.  So that would indeed be 
> informatically difficult to represent in signed 32-bit arithmetic.  Perhaps 
> when this plant is assembled, the researchers will need to artificially 
> subdivide the chromosomes in some way, or the formats will have moved on by 
> then.
>
>> [1] 
>> https://docs.google.com/spreadsheet/ccc?key=0Aqs5UFlky_s6dDlmVVFib3FoSm1tS1JIQzMxY0RVb2c
>
> It appears that perhaps there is in fact a convention of subdividing large 
> plant chromosomes into long and short arms for reference purposes: your 
> spreadsheet lists e.g. 2BL and 2BS separately.
>
> One of the samtools bug reports about large chromosomes was about barley [1], 
> with seven chromosomes 1H...7H.  I downloaded Ensembl's barley genome while 
> attempting to reproduce it, but was thoroughly confused by the apparently 
> twenty chromosomes there: see 
> http://plants.ensembl.org/Hordeum_vulgare/Location/Genome which seems to have 
> mostly individual chromosome arms in 2HL, 2HS etc but also enormous 1-7 
> chromosomes.  Dan, I think you just volunteered to explain what on earth is 
> going on here! :-)
>
>     John
>
>
>> On 18 October 2013 11:50, John Marshall <[email protected]> wrote:
>>> [snip]
>>> Hopefully 2 Gbp suffices for most species that people are interested in.  
>>> In principle the existing file formats could be compatibly pushed to 4 Gbp, 
>>> but this would require great care with getting signed v. unsigned 
>>> arithmetic correct in the implementations so would likely not really be 
>>> worthwhile.
>
> [1] https://github.com/samtools/samtools/issues/116
>
> --
>  The Wellcome Trust Sanger Institute is operated by Genome Research
>  Limited, a charity registered in England with number 1021457 and a
>  company registered in England with number 2742969, whose registered
>  office is 215 Euston Road, London, NW1 2BE.

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