That's interesting. I wouldn't have thought there was any controversy, and said it was pretty obvious that bats flew before the echolocated. Echolocation would be of limited use if you weren't flying, and anyway a major subdivision of bats, the megabats (family Pteropodidae, flying foxes) don't echolocate, although they fly. I guess that just shows how little I know (or how desperate specialists are for controversies to write papers about). -- Mixon
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