The mobile version of Firefox and upcoming versions will play H264/AAC/MP3, but will use the system codecs if installed. Since Firefox does not bundle the codecs, they should be fine right? Rumor is the GNU/Linux versions will use Gstreamer's plugins for H264 playback and the Windows versions use the built in codecs of Windows 7.

Chromium uses FFMPEG and I really like how it gives you a choice between royalty free only (chromium-codecs-ffmpeg) or not (chromium-codecs-ffmpeg-extra): http://packages.ubuntu.com/source/precise-updates/chromium-browser

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