I would not use smtp-auth because my mail server is simply relaying a mail to them.

I think I noticed this happening when my mail server sends mail to ext-mx01.kolabsys.com, because mail is accepted if sent via ext-mx02.kolabsys.com.

Here are the headers from the email that was bounced, and here we see the From set to users@lists.roundcube.net upon reply, and now I know exactly why this happens! I have a Roundcube plugin installed called custom_from.
     https://github.com/r3c/Custom-From
"When replying to an e-mail sent to you through an address not in your identities list, plugin will automatically fire and set "From:" header to the address the original e-mail was sent to."


MIME-Version: 1.0
Content-Type: text/plain; charset=UTF-8;
 format=flowed
Content-Transfer-Encoding: 7bit
Date: Wed, 27 Nov 2013 13:23:56 +0100
From: Roundcube Users mailing list <users@lists.roundcube.net>
To: users@lists.roundcube.net
X-Sender: users@lists.roundcube.net
User-Agent: Roundcube Webmail/0.9.5


This explains everything :)



On 2013-11-27 14:54, Benny Pedersen wrote:
Simon Loewenthal skrev den 2013-11-27 14:22:

said: 553 5.7.1 <users@lists.roundcube.net>: Sender address rejected: not logged in

the above error is that you either use there domain or not using smtp auth

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