Thanks for all who answered. Changing the "official hostname" of the primary domain to match solved the problem... So apparently they required the HELO hostname to match exactly.
Remember, that may have solved the problem, but it also shows that they are using a "crazy anti-spam test". What that means is that since you have jumped through hoops to get your mail to them, you may have to jump back through those same hoops to get mail to someone else. In general, when someone runs a super extreme spam test like that, it is best not to give in to them.
-Scott
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