For anyone else struggling with this problem, the behaviour pattern of USB lock-ups has completely changed when I swapped the Pi power supply for a different one. My system was locking up at 1 to 2 day intervals on the old power supply; I am now at 10 days without a problem on the new power supply. It seems that the USB interface on the WH1080 is voltage (or current) sensitive. This would be consistent with the recommendation to use a powered USB hub.
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