Your condition will always influence your ride quality.   Sometimes just 
the lack of coffee and a cruddy day at work really sets you wrong.  
Luckily, a ride usually fixes that.

Grant's designs always whisper in your ear to take the longer way home, 
then less direct route, the more adventurous vector.  Always.

There are a number of reasons for that, with the two main being the ability 
to run high quality, larger volume tires and the position and geometry of 
the bike.  As Steve points out, there's nothing inherently "non-modern" 
about either of those concepts.

And, honestly, you could work in carbon fiber or aluminum or titanium or 
thermoplastic and retain some of those attributes.  Steel's specific 
attributes have other benefits in addition to the ductility that allows 
flex.  

On long rides, vibration is a killer.  Damping that vibration is done by 
the tires, wheels, frame and rider in that order.  Thin tires, rigid 
wheels, stiff frame all mean the rider will take up more of the effort.    
If you combine that with a race-oriented "aero" position - bars well below 
the saddle so that you have a horizontal back, you force the arms and hips 
to act as bulwarks in the bridge that is your backbone.  More upright (with 
significant physiological variability, of course) allows your body's 
natural "spring" system (spinal curve, knees, elbows) to work the way they 
evolved.  

All of that means you ask less of your body with each mile. 

My last open-wheeled racer wasn't particularly aggressive, but  at ~60 
miles, I was in need of a break, even during my highest mileage years. It 
was mostly that my body didn't want to come up out of the position it had 
been holding.  I was fine when I was riding.   The bike handled well, so it 
was never a need of trying to keep on top of it as I got more tired. 

I've done centuries and 200K brevets on both the Hilsen and the Quickbeam.  
Some where extremely taxing, but that definitely was an engine issue. 

It's pretty simple in my book - Start Comfortable, Stay Comfortable. 

- J

cyclofiend.com / about.me/cyclofiend

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