I did a couple of tours on this restored long chain stay Trek 728: http://cyclofiend.com/cc/2008/cc484-joelmatthews0508.html before donating to a local bike co-op. The ride with the 35 Schwalbes was very comfortable. Never tried it with narrower tires. The 728 had a mid to high trail fork. The only issue I had with the bike is steering was somewhat lazy in the tight corners. Perhaps a builder would have some ideas how this could have been corrected. As it is, Trek dropped the 728 the next year without tweaking the design.
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